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spectacle-pod

Palmer's spectaclepod

Habit Annuals, biennials, or, rarely, perennials; not scapose; densely pubescent throughout, trichomes usually subsessile and stellate, mixed with minutely stalked, dendritic ones, rarely unbranched. Annuals or biennials.
Stems

erect [ascending], unbranched or branched proximally, branched distally.

unbranched proximally, branched distally, (3–)4–8(–10) dm.

Leaves

basal and cauline;

petiolate or sessile;

basal not rosulate, petiolate, blade margins dentate, pinnatifid, or lobed;

cauline sessile, subsessile, or shortly petiolate, blade (base not auriculate), margins entire or lobed.

Basal leaves

petiole 1–4(–6) cm;

blade lanceolate to oblong or ovate, (2–)4–8(–10) cm × (10–)15–25(–40) mm, base cuneate to obtuse, margins dentate.

Cauline leaves

(distal) sessile;

blade usually ovate to narrowly oblong, rarely lanceolate, base obtuse to truncate, margins entire, sometimes repand.

Racemes

(corymbose), considerably elongated in fruit.

Flowers

sepals widely spreading to reflexed, oblong, (equal), lateral pair not saccate basally;

petals (spreading), white or lavender, obovate (longer than sepals), claw well-differentiated from blade, (apex rounded);

stamens (somewhat spreading), tetradynamous;

filaments not dilated basally;

anthers oblong [sagittate];

nectar glands distinct, lateral and median present; (gynophore to 0.5 mm).

sepals 3.5–5 × 1–1.5 mm, pubescent abaxially;

petals white or lavender, (7–)8–10(–12) × 4–6(–7) mm, attenuate to claw, claw 2–3 mm, not expanded basally;

filaments lavender or white, 3–4 mm;

anthers 1–1.5 mm.

Fruiting pedicels

divaricate, sometimes ascending or slightly reflexed, slender.

divaricate, (8–)10–16(–20) mm.

Fruits

silicles, subsessile, breaking into two 1-seeded units at maturity, didymous, suborbicular, or broader than long, winged, strongly angustiseptate;

valves keeled, enclosing seeds when falling off, not winged, narrowly winged, or margined, indurated around margin, glabrous or pubescent;

replum concealed by valve margin;

septum complete (ca. 0.1 mm wide);

ovules 2 per ovary;

style relatively short or obsolete, (terete or flattened);

stigma conical, decurrently 2-lobed (appearing entire).

each valve suborbicular or orbicular, (7–)8–10 × (6–)7–10 mm, base rounded, apex truncate, with or without narrow margin beyond indurated part surrounding seeds, glabrous or pubescent;

style (0.3–)0.6–1(–1.2) mm.

Seeds

aseriate, strongly flattened, not winged, broadly oblong to suborbicular;

seed coat (smooth), not mucilaginous when wetted;

cotyledons accumbent to obliquely incumbent.

suborbicular-ovoid, 3–4 × 2.5–3 mm.

x

= 9.

2n

= 18.

Dimorphocarpa

Dimorphocarpa candicans

Phenology Flowering Apr–Sep.
Habitat Sandy hills and plains, prairies, sand dunes
Elevation 100-800 m (300-2600 ft)
Distribution
from USDA
sw United States; n Mexico
[BONAP county map]
from FNA
KS; NM; OK; TX
[BONAP county map]
Discussion

Species 4 (3 in the flora).

Dimorphocarpa membranacea (Payson) Rollins is known from northern Mexico (San Luis Potosí, Tamaulipas).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves (7-)8-10 mm; petals (7-)8-10(-12) mm, claws not expanded basally; distal cauline leaves sessile, blades usually ovate to narrowly oblong, rarely lanceolate, bases obtuse to truncate.
D. candicans
1. Fruit valves 4-5.5(-6.5) mm; petals (3.5-)4-7(-8) mm, claws expanded basally; distal cauline leaves shortly petiolate or subsessile, blades linear to lanceolate, bases cuneate
→ 2
2. Distal cauline blades: margins pinnately lobed; fruit valves nearly as long as wide, rounded apically.
D. pinnatifida
2. Distal cauline blades: margins usually entire, rarely dentate; fruit valves often longer than wide, truncate apically.
D. wislizeni
Source FNA vol. 7, p. 604. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 605.
Parent taxa Brassicaceae > tribe Physarieae Brassicaceae > tribe Physarieae > Dimorphocarpa
Sibling taxa
D. pinnatifida, D. wislizeni
Subordinate taxa
D. candicans, D. pinnatifida, D. wislizeni
Synonyms Iberis candicans, D. palmeri, Dithyrea wislizeni var. palmeri
Name authority Rollins: Publ. Bussey Inst. Harvard Univ. 1979: 20, plates 3, 4. (1979) (Rafinesque) Rollins: Cruciferae Continental N. Amer., 361. (1993)
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