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Habit Plants usually blackish green. Plants green to tan.
Stem(s)

leaves spreading-incurved and twisted to incurved-appressed when dry, spreading to spreading-recurved and not keeled when moist, monomorphic, short-lanceolate or sometimes short-ovate to long-elliptic, broadly concave adaxially across leaf, usually 1–2.5 mm, base scarcely differentiated in shape to ovate, margins usually recurved at mid leaf, entire, apex broadly acute or occasionally narrowly so, not fragile;

costa percurrent or ending a few cells below the apex, broader at mid leaf, occasionally weakly spurred, with a low adaxial pad of cells, adaxial costal cells quadrate, 4–6 cells wide at mid leaf, guide cells in 1–2 layers;

basal laminal cells differentiated medially or across leaf, quadrate to short-rectangular, walls very thin and not perforated;

distal laminal cells 7–12 wide, 1:1 or occasionally transversely elongate below, papillae usually distinct, low or simple to 2-fid, occasionally absent, lumens oval to rounded-quadrate, walls thin to evenly thickened, convex on both sides of lamina, 2-stratose in one or more rows along margins.

central strand and sclerodermis usually present, hyalodermis rare.

Leaves

usually broadly lanceolate to narrowly elliptical, apex blunt to acute, margins recurved basally;

distal laminal cells if 2-stratose stacked directly over one another, walls smooth or papillose, KOH yellow or red;

costa with or without a differentiated abaxial epidermis;

gemmae occasionally present, usually clavate.

Seta

0.7–1 cm.

Capsule

1–1.9 mm;

peristome teeth 32, linear, weakly twisted, to 600 µm, occasionally rudimentary.

Spores

11–15 µm.

Specialized

asexual reproduction by multicellular tubers on proximal rhizoids.

Distal

laminal KOH reaction variously negative or yellow- or orange- or red-brown.

Peristome

usually present.

Didymodon australasiae

Pottiaceae subfam. barbuloideae

Phenology Capsules mature winter–spring.
Habitat Soil, gypsum, acid rock, ledges, sandstone, silt
Elevation moderate to high elevations (300-2000 m) (moderate to high elevations (1000-6600 ft))
Distribution
from FNA
AZ; CA; CO; NM; NV; OR; TX; Mexico; Central America; South America; Europe; n Africa; s Africa; Pacific Islands (New Zealand); Australia
Nearly worldwide
Discussion

In North American Didymodon, hydroids are found in the costae of D. anserinocapitatus, D. australasiae, D. nevadensis, D. norrisii, D. umbrosus, D. revolutus, and D. vinealis, and in these species the adaxial stereid band is usually absent in well developed specimens. Intergrades exist between D. australasiae and D. umbrosus, but the extreme forms are common and quite distinctive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 33 (16 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 547. FNA vol. 27, p. 519.
Parent taxa Pottiaceae > subfam. Barbuloideae > Didymodon Pottiaceae
Sibling taxa
D. anserinocapitatus, D. asperifolius, D. bistratosus, D. brachyphyllus, D. eckeliae, D. fallax, D. ferrugineus, D. giganteus, D. johansenii, D. leskeoides, D. maschalogena, D. maximus, D. murrayae, D. nevadensis, D. nicholsonii, D. nigrescens, D. norrisii, D. perobtusus, D. revolutus, D. rigidulus, D. subandreaeoides, D. tectorum, D. tophaceus, D. umbrosus, D. vinealis
Subordinate taxa
Synonyms Tortula australasiae, D. diaphanobasis, Husnotiella torquescens, Trichostomopsis australasiae, Trichostomopsis brevifolia, Trichostomopsis diaphanobasis, Trichostomopsis fayae family Pottiaceae tribe Barbuleae
Name authority (Hooker & Greville) R. H. Zander: Phytologia 41: 21. (1978) (Herzog) Hilpert: Beih. Bot. Centralbl. 20(2): 612. (1933)
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