The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

black beard moss, black didymodon moss

Habit Plants usually blackish green. Plants usually red- to black-brown, occasionally yellow- or orange-brown at apex.
Stem(s)

leaves spreading-incurved and twisted to incurved-appressed when dry, spreading to spreading-recurved and not keeled when moist, monomorphic, short-lanceolate or sometimes short-ovate to long-elliptic, broadly concave adaxially across leaf, usually 1–2.5 mm, base scarcely differentiated in shape to ovate, margins usually recurved at mid leaf, entire, apex broadly acute or occasionally narrowly so, not fragile;

costa percurrent or ending a few cells below the apex, broader at mid leaf, occasionally weakly spurred, with a low adaxial pad of cells, adaxial costal cells quadrate, 4–6 cells wide at mid leaf, guide cells in 1–2 layers;

basal laminal cells differentiated medially or across leaf, quadrate to short-rectangular, walls very thin and not perforated;

distal laminal cells 7–12 wide, 1:1 or occasionally transversely elongate below, papillae usually distinct, low or simple to 2-fid, occasionally absent, lumens oval to rounded-quadrate, walls thin to evenly thickened, convex on both sides of lamina, 2-stratose in one or more rows along margins.

leaves appressed when dry, spreading and not keeled when moist, monomorphic, long-ovate to lanceolate, broadly concave adaxially across leaf, usually 0.8–1.7 mm, base weakly differentiated in shape, long-ovate, margins usually broadly recurved to revolute to mid leaf or to near apex, minutely crenulate, apex acute to narrowly acuminate, often weakly cucullate;

costa percurrent or ending 2–4 cells below the apex, little tapering, not strongly spurred, without an adaxial pad of cells, adaxial costal cells rectangular, 2 cells wide at mid leaf grading to 4 below, guide cells in 1 layer;

basal laminal cells differentiated medially, walls thick, rectangular, not perforated;

distal laminal cells in rows, 7–11(–14) µm wide, 1:1 or often longitudinally elongate, papillae apparently absent but visible in section as low, flattened to multiplex capitulate lenses, 1–3 per lumen, lumens ovate, walls evenly thickened and weakly convex on both sides of lamina, 1-stratose.

Seta

0.7–1 cm.

0.7–0.9 cm.

Capsule

1–1.9 mm;

peristome teeth 32, linear, weakly twisted, to 600 µm, occasionally rudimentary.

1.5–2.8 mm;

peristome teeth 32, linear, straight to twisted 1.5 times, 100–600 µm.

Spores

11–15 µm.

9–13 µm.

Specialized

asexual reproduction by multicellular tubers on proximal rhizoids.

asexual reproduction absent.

Distal

laminal KOH reaction variously negative or yellow- or orange- or red-brown.

laminal KOH reaction red.

Didymodon australasiae

Didymodon nigrescens

Phenology Capsules mature winter–spring. Capsules mature spring–fall.
Habitat Soil, gypsum, acid rock, ledges, sandstone, silt Limestone, frostboils, outcrops, cliff faces, often near streams and waterfalls
Elevation moderate to high elevations (300-2000 m) (moderate to high elevations (1000-6600 ft)) low to moderate elevations (0-700 m) (low to moderate elevations (0-2300 ft))
Distribution
from FNA
AZ; CA; CO; NM; NV; OR; TX; Mexico; Central America; South America; Europe; n Africa; s Africa; Pacific Islands (New Zealand); Australia
from FNA
AK; BC; NT; Central America (Guatemala); e Asia
Discussion

In North American Didymodon, hydroids are found in the costae of D. anserinocapitatus, D. australasiae, D. nevadensis, D. norrisii, D. umbrosus, D. revolutus, and D. vinealis, and in these species the adaxial stereid band is usually absent in well developed specimens. Intergrades exist between D. australasiae and D. umbrosus, but the extreme forms are common and quite distinctive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The specimens cited as Didymodon nigrescens from the Firth River Basin by W. C. Steere (1978), determined by R. H. Zander, are actually Didymodon subandreaeoides. The distinguishing characters of D. nigrescens are the blackish coloration when dry (red in KOH), thin costa and distal laminal margins minutely crenulate by the small, bulging marginal cells. This and the following two species appear to be closely related by the rather distinctive crenulation of the leaf apices. Didymodon asperifolius is similar but may be distinguished by its longer leaves, straight or reflexed to strongly recurved when wet, distal margins recurved and smooth, distal laminal cells larger, 10–13 µm wide, epapillose or papillae simple, costa usually rather wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 547. FNA vol. 27, p. 549.
Parent taxa Pottiaceae > subfam. Barbuloideae > Didymodon Pottiaceae > subfam. Barbuloideae > Didymodon
Sibling taxa
D. anserinocapitatus, D. asperifolius, D. bistratosus, D. brachyphyllus, D. eckeliae, D. fallax, D. ferrugineus, D. giganteus, D. johansenii, D. leskeoides, D. maschalogena, D. maximus, D. murrayae, D. nevadensis, D. nicholsonii, D. nigrescens, D. norrisii, D. perobtusus, D. revolutus, D. rigidulus, D. subandreaeoides, D. tectorum, D. tophaceus, D. umbrosus, D. vinealis
D. anserinocapitatus, D. asperifolius, D. australasiae, D. bistratosus, D. brachyphyllus, D. eckeliae, D. fallax, D. ferrugineus, D. giganteus, D. johansenii, D. leskeoides, D. maschalogena, D. maximus, D. murrayae, D. nevadensis, D. nicholsonii, D. norrisii, D. perobtusus, D. revolutus, D. rigidulus, D. subandreaeoides, D. tectorum, D. tophaceus, D. umbrosus, D. vinealis
Synonyms Tortula australasiae, D. diaphanobasis, Husnotiella torquescens, Trichostomopsis australasiae, Trichostomopsis brevifolia, Trichostomopsis diaphanobasis, Trichostomopsis fayae Barbula nigrescens, Barbula rufofusca
Name authority (Hooker & Greville) R. H. Zander: Phytologia 41: 21. (1978) (Mitten) K. Saito: J. Hattori Bot. Lab. 39: 510. (1975)
Web links