Flora of North America species comparison

leaves spreading-incurved and twisted to incurved-appressed when dry, spreading to spreading-recurved and not keeled when moist, monomorphic, short-lanceolate or sometimes short-ovate to long-elliptic, broadly concave adaxially across leaf, usually 1–2.5 mm, base scarcely differentiated in shape to ovate, margins usually recurved at mid leaf, entire, apex broadly acute or occasionally narrowly so, not fragile;

costa percurrent or ending a few cells below the apex, broader at mid leaf, occasionally weakly spurred, with a low adaxial pad of cells, adaxial costal cells quadrate, 4–6 cells wide at mid leaf, guide cells in 1–2 layers;

basal laminal cells differentiated medially or across leaf, quadrate to short-rectangular, walls very thin and not perforated;

distal laminal cells 7–12 wide, 1:1 or occasionally transversely elongate below, papillae usually distinct, low or simple to 2-fid, occasionally absent, lumens oval to rounded-quadrate, walls thin to evenly thickened, convex on both sides of lamina, 2-stratose in one or more rows along margins.

leaves appressed and incurved when dry, patent to spreading-recurved and not keeled when moist, monomorphic, long-lanceolate, grooved adaxially along the costa, especially near leaf apex, 2–3 mm, base scarcely differentiated or short-oblong, margins narrowly recurved in proximal 1/3–1/2 of leaf, often broadly crenate (ca. 8–10 cells across), sinuose or weakly notched in distal 2/3–3/4 of leaf, apex narrowly acute or narrowly blunt and long-acuminate, somewhat fragile;

costa percurrent or short-excurrent, not strongly spurred, not much widened or tapering, lacking an adaxial pad of cells, adaxial costal cells quadrate, ca. 4 cells wide at mid leaf, guide cells in 2 layers;

basal laminal cells very weakly differentiated medially, walls thin to weakly thickened, quadrate to short-rectangular, not perforated;

distal laminal cells 7–9 µm wide, 1:1, nearly smooth or papillae simple or 2-fid, 2–3 per lumen, lumens rounded-quadrate, walls thin or evenly thickened, weakly convex on both sides of lamina, distal 2/3–3/4 of the leaf margins 2-stratose with 1–2 rows of cells similar to the medial cells of the lamina.

0.7–1 cm.

0.8–1.2 cm.

apparently dioicous;

perigonia not seen;

perichaetial leaves little different from the cauline or with wider bases.

1–1.9 mm;

peristome teeth 32, linear, weakly twisted, to 600 µm, occasionally rudimentary.

2–2.4 mm;

peristome teeth 32, linear, weakly twisted or to 1/2 times, ca. 700–1000 µm. Spores 8–10(–13) µm. Distal laminal KOH reaction yellow-red.

11–15 µm. Distal laminal KOH reaction variously negative or yellow- or orange- or red-brown.

asexual reproduction by multicellular tubers on proximal rhizoids.

asexual reproduction apparently by fragile leaves.