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Habit Plants usually blackish green.

leaves spreading-incurved and twisted to incurved-appressed when dry, spreading to spreading-recurved and not keeled when moist, monomorphic, short-lanceolate or sometimes short-ovate to long-elliptic, broadly concave adaxially across leaf, usually 1–2.5 mm, base scarcely differentiated in shape to ovate, margins usually recurved at mid leaf, entire, apex broadly acute or occasionally narrowly so, not fragile;

costa percurrent or ending a few cells below the apex, broader at mid leaf, occasionally weakly spurred, with a low adaxial pad of cells, adaxial costal cells quadrate, 4–6 cells wide at mid leaf, guide cells in 1–2 layers;

basal laminal cells differentiated medially or across leaf, quadrate to short-rectangular, walls very thin and not perforated;

distal laminal cells 7–12 wide, 1:1 or occasionally transversely elongate below, papillae usually distinct, low or simple to 2-fid, occasionally absent, lumens oval to rounded-quadrate, walls thin to evenly thickened, convex on both sides of lamina, 2-stratose in one or more rows along margins.


0.7–1 cm.


1–1.9 mm;

peristome teeth 32, linear, weakly twisted, to 600 µm, occasionally rudimentary.


11–15 µm.


asexual reproduction by multicellular tubers on proximal rhizoids.


laminal KOH reaction variously negative or yellow- or orange- or red-brown.

Didymodon australasiae

Phenology Capsules mature winter–spring.
Habitat Soil, gypsum, acid rock, ledges, sandstone, silt
Elevation moderate to high elevations (300-2000 m) (moderate to high elevations (1000-6600 ft))
from FNA
AZ; CA; CO; NM; NV; OR; TX; Mexico; Central America; South America; Europe; n Africa; s Africa; Pacific Islands (New Zealand); Australia

In North American Didymodon, hydroids are found in the costae of D. anserinocapitatus, D. australasiae, D. nevadensis, D. norrisii, D. umbrosus, D. revolutus, and D. vinealis, and in these species the adaxial stereid band is usually absent in well developed specimens. Intergrades exist between D. australasiae and D. umbrosus, but the extreme forms are common and quite distinctive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 547.
Parent taxa Pottiaceae > subfam. Barbuloideae > Didymodon
Sibling taxa
D. anserinocapitatus, D. asperifolius, D. bistratosus, D. brachyphyllus, D. eckeliae, D. fallax, D. ferrugineus, D. giganteus, D. johansenii, D. leskeoides, D. maschalogena, D. maximus, D. murrayae, D. nevadensis, D. nicholsonii, D. nigrescens, D. norrisii, D. perobtusus, D. revolutus, D. rigidulus, D. subandreaeoides, D. tectorum, D. tophaceus, D. umbrosus, D. vinealis
Synonyms Tortula australasiae, D. diaphanobasis, Husnotiella torquescens, Trichostomopsis australasiae, Trichostomopsis brevifolia, Trichostomopsis diaphanobasis, Trichostomopsis fayae
Name authority (Hooker & Greville) R. H. Zander: Phytologia 41: 21. (1978)
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