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dicranum moss

boulder broom moss, dicranum moss

Habit Plants in loose tufts, green to yellowish green or yellowish brown, dull. Plants in loose tufts, dark green above, blackish green below, dull.
Stems

2–11 cm, densely tomentose below with reddish brown rhizoids, interruptedly foliate, the leaves crowded in some parts forming swollen, globose regions.

1.5–3.5 cm, sparsely tomentose with light- to reddish brown rhizoids.

Leaves

erect-spreading, arched and loosely imbricate with contorted apices when dry, ± undulate to rugose, (2.5–)5–6.5(–9) × 1–1.7 mm, concave below, tubulose above, acute, ovate at base, widest point at or somewhat below middle;

margins serrulate to serrate in distal half, plane to involute;

laminae 1-stratose or with 2-stratose regions near costa and on margins;

costa percurrent to shortly excurrent, 1/8–1/4 the width of the leaves at base, serrulate or toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two stereid bands extending to apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells weakly to strongly bulging;

leaf cells strongly papillose above on abaxial surface, sometimes a few papillae on adaxial surface;

alar cells 2-stratose, distinctly differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (30–)49–68(–101) × (5–)10–11(–15) µm, abruptly shorter in distal half of leaf;

distal laminal cells short, not or indistinctly pitted, cell walls irregularly thickened, (7–)11–12(–23) × (5–)7–13(–20) µm. Sexual condition pseudomonoicous;

dwarf males on rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

erect-spreading, some weakly falcate-secund, crisped when dry, smooth, (3–)4–5(–7) × 0.5–0.8 mm, some leaf tips broken off, concave below, subtubulose above, narrowly lanceolate to a long, narrow subula occupied mainly by the excurrent costa, apex acute;

margins serrate to serrulate in distal half;

laminae 1- or 2-stratose above, sometimes 2-stratose in patches;

costa excurrent, 1/4–1/3 the width of the leaves at base, abaxially toothed or papillose in distal half, abaxial ridges absent, with a row of guide cells, two stereid bands not extending above the leaf middle, adaxial and abaxial epidermal layers of cells somewhat differentiated or with a few cells enlarged in both layers, cell walls between lamina cells strongly bulging in distal part of leaf lamina;

leaf cells smooth or abaxially prorate above;

alar cells mostly 1-stratose or sometimes with a few 2-stratose cells, distinctly differentiated, often extending to costa;

proximal laminal cells elongate-rectangular, not pitted or with few pits, (19–)24–33(–42) × (4)5–6(9) µm;

distal laminal cells regularly quadrate to short-rectangular, not pitted, (5–)10–11(–16) × (4)5–6(9) µm. Sexual condition dioicous;

male plants as tall as female plants but usually more slender;

interior perichaetial leaves abruptly long-acuminate, convolute-sheathing.

Seta

1–3 cm, solitary, reddish yellow or brown.

1–2 cm., solitary, brown or reddish brown.

Capsule

1.5–2.5 mm, arcuate, inclined to horizontal, contracted below mouth and furrowed when dry, yellowish brown to brown;

operculum 1.5–2.5 mm.

1.5–3 mm, straight and erect, smooth, slightly furrowed when dry, reddish brown;

operculum 1.5–2 mm.

Spores

16–24 µm.

14–28 µm.

Dicranum spurium

Dicranum fulvum

Phenology Capsules mature spring. Capsules mature in spring.
Habitat Sandy soil, decayed logs, acidic rock, humus over exposed bluffs or outcrops, in pine woods, sometimes bogs Sandstone, limestone or acidic rock outcrops, bluffs, cliff faces, and boulders in deciduous woodlands, especially along streams, rarely on bases of tree trunks, fallen trees and logs
Elevation 0-1400 m (0-4600 ft) 200-1900 m (700-6200 ft)
Distribution
from FNA
AR; CT; GA; IL; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; OK; PA; TN; VA; VT; WV; NF; NS; ON; PE; QC; Europe
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from FNA
AL; AR; CT; DC; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WI; WV; NB; NS; ON; QC; Europe; Asia
[WildflowerSearch map]
Discussion

Dicranum spurium is easily recognized by the turgid aspect of the plants with arched, imbricate leaves and with interruptedly foliate stems, i.e., some regions that appear swollen or globose as a result of several growth periods. It is likely to be confused only with 11. D. condensatum (see discussion thereunder).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum fulvum has dull, blackish green plants, especially evident from the leaves on the basal part of the stems, leaves crisped when dry, broad costae that cover 1/4–1/3 the width of the leaves at base, laminae that are mostly 2-stratose in the distal half of the leaf, and erect, cylindrical capsules that are slightly furrowed when dry. It is sometimes confused with D. fuscescens but that species has keeled leaves in the distal half, with 2-stratose regions restricted to the margins, and horizontal capsules, whereas D. fulvum has subtubulose leaves, with almost entirely 2-stratose laminae above, and erect capsules. When D. fulvum has some of its leaf apices broken off, which is not uncommon, it can be confused with D. viride, which has the majority of its leaf tips absent. The usual occurrence on rock of plants of D. fulvum as opposed to the usual occurrence on bases of tree trunks of D. viride often gives some clue to their identity. Morphologically, the proximal leaf cells of the two differ: D. fulvum has shorter cells, averaging 24–33 µm, compared to the longer cells of D. viride, averaging 33–42 µm.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 417.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
Name authority Hedwig: Sp. Musc. Frond., 141. (1801) Hooker: Musci Exot. 2: plate 149. (1819)
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