Dicranum polysetum |
Dicranum viride |
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dicranum moss, wavy broom moss, wavy-leaf heron's-bill moss, wavy-leaf moss |
dicranum moss |
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Habit | Plants in loose tufts, light green, glossy. | Plants in loose to dense tufts, yellowish to dark green, glossy to dull. |
Stems | 4–15 cm, densely tomentose with whitish or reddish rhizoids. |
1.5–4.5 cm, sparsely tomentose with whitish to reddish brown rhizoids. |
Leaves | erect to spreading, ± flexuose, little changed when dry, strongly undulate, (5.5–)7–9.5(–10.5) × 1–2 mm, lanceolate, concave proximally, keeled above, acute; margins strongly toothed in the distal half; laminae 1-stratose; costa ending before apex, occupying ca. 1/16–1/8 of leaf base, strong, with two toothed ridges distally on abaxial surface, with one row of guide cells, two stereid bands, adaxial epidermal layer of cells not differentiated, abaxial layer with a few cells enlarged; cell walls between lamina cells not bulging; leaf cells smooth; alar cells 2-stratose, well-differentiated, not extending to costa; proximal laminal cells elongate, pitted, (45–)78–129(–156) × (5–)9–10(–14) µm; distal laminal cells shorter, sinuose, pitted, (42–)64–68(–115) × (4–)9–10(–13)µm. Sexual condition pseudomonoicous; dwarf male plants on stem rhizoids of female plants; interior perichaetial leaves abruptly long-acuminate, convolute-sheathing. |
erect-spreading or somewhat falcate-secund, rigid, erect below and flexuose above when dry, smooth, (3–)4–6(–7) × 0.5–0.8 mm, most of the leaf tips broken off, concave below to canaliculate above, from a lanceolate base to a long-acuminate, acute apex (when present); margins entire, sometimes slightly denticulate at apex; laminae 1-stratose or with some 2-stratose regions in patches between margin and costa; costa excurrent, 1/5–1/4 the width of the leaves at base, smooth or slightly rough on abaxial surface in distal half, abaxial ridges absent, with a row of guide cells, two weakly developed stereid bands above and below, not extending above the leaf middle, adaxial and abaxial epidermal layers of cells not differentiated or with a few cells enlarged in both layers; cell walls between lamina cells not or weakly bulging; leaf cells usually smooth; alar cells 1-stratose or with a few 2-stratose regions, well-differentiated, often extending to costa; proximal laminal cells rectangular to short-rectangular, not pitted or with few pits, (11–)33–42(–51) × (7–)9–10(–13) µm; median laminal cells regularly quadrate, not pitted, (9–)15–22(–26) × (5–)7–8(–11) µm; distal laminal cells small, quadrate, not pitted. |
Seta | 1.5–4 cm, usually aggregate, 3–6 per perichaetium, brown or reddish brown. |
1–1.6 cm, solitary, yellow to reddish brown. |
Sexual condition | dioicous; male plants as large as females; interior perichaetial leaves abruptly long-acuminate, convolute-sheathing. |
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Capsule | 2–3.5 mm, arcuate, inclined to horizontal, furrowed when dry, yellowish brown or reddish brown; operculum 2–4 mm. |
1.5–2.5 mm, straight, erect, smooth, slightly furrowed when dry, brown to reddish brown; operculum 1–1.5 mm. |
Spores | 12–24 µm. |
9–22 µm. |
Dicranum polysetum |
Dicranum viride |
|
Phenology | Capsules mature spring. | Capsules mature in spring. |
Habitat | Commonly on humus, soil over acidic or calcareous rock, and decaying wood in deciduous or more often coniferous forests, occasionally in bogs, fens, and swamps | Commonly growing on the base of trees (usually deciduous but sometimes coniferous, especially Thuja), rotten logs, stumps, rarely soil and acidic or limestone rock |
Elevation | 10-2100 m (0-6900 ft) | 40-1400 m (100-4600 ft) |
Distribution |
AK; CO; CT; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; PE; QC; SK; YT; Europe; Asia
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AK; IA; IL; IN; KY; MA; MD; ME; MI; MN; NC; NH; NY; OH; PA; TN; VA; VT; WI; WV; MB; NB; NF; NS; ON; PE; QC; Europe; Asia |
Discussion | W. L. Peterson (1979) recorded Dicranum polysetum from Nunavut. It is one of the largest, most conspicuous, and most easily recognized species in the genus in North America. The species is immediately recognized by the large stems, 4–15 cm, with a dense mat of whitish or reddish tomentum, giving the stems a thickened appearance, by the shiny, strongly undulate, wide-spreading leaves with margins strongly toothed in the distal half and by the clustered sporophytes (3–6 per perichaetium). Microscopically, the species is readily distinguished by the elongate, abundantly pitted cells throughout the leaf and by the toothed ridges on the abaxial surface of the costa in the distal part of the leaf. Dicranum polysetum is one of the few species in North America that has no apparent intergradations with any other species of the genus. It is common in southeastern Canada, northeastern United States, and the Canadian Rocky Mountains, where it extends south to Wyoming and Colorado, occurring in both states in a small number of localities. It is rare west of the Rockies, where it is known from only a few localities in northwestern Montana, southern Idaho and northeastern Washington and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Dicranum viride has been reported from Alaska by I. A. Worley and Z. Iwatsuki (1970) and from Kentucky by J. A. Snider et al. (1988). It is a not uncommon species of the eastern United States and southeastern Canada. It rarely produces sporophytes and is distinctive because of the fragile, deciduous leaf tips, which probably serve as a means of asexual reproduction. It has on occasion been confused with 22. D. fulvum, which has nearly the same distribution, and rarely with 21. D. fragilifolium where their ranges overlap in the Great Lakes region. For distinctions see the discussions under those species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 401. | FNA vol. 27, p. 417. |
Parent taxa | Dicranaceae > Dicranum | Dicranaceae > Dicranum |
Sibling taxa | ||
Synonyms | D. rugosum | Campylopus viridis, D. fulvum subsp. viride, D. fulvum var. viride |
Name authority | Swartz: Monthly Rev., ser. 2, 34: 538. (1801) | (Sullivant & Lesquereux) Lindberg: Hedwigia 2: 70. (1863) |
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