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dicranum moss, wavy broom moss, wavy-leaf heron's-bill moss, wavy-leaf moss

broken-leaf heron's-bill moss, broken-leaf moss, dicranum moss

Habit Plants in loose tufts, light green, glossy. Plants in dense tufts, light green to yellowish green, glossy.
Stems

4–15 cm, densely tomentose with whitish or reddish rhizoids.

0.5–3 cm, tomentose below with whitish or reddish brown rhizoids.

Leaves

erect to spreading, ± flexuose, little changed when dry, strongly undulate, (5.5–)7–9.5(–10.5) × 1–2 mm, lanceolate, concave proximally, keeled above, acute;

margins strongly toothed in the distal half;

laminae 1-stratose;

costa ending before apex, occupying ca. 1/16–1/8 of leaf base, strong, with two toothed ridges distally on abaxial surface, with one row of guide cells, two stereid bands, adaxial epidermal layer of cells not differentiated, abaxial layer with a few cells enlarged;

cell walls between lamina cells not bulging;

leaf cells smooth;

alar cells 2-stratose, well-differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (45–)78–129(–156) × (5–)9–10(–14) µm;

distal laminal cells shorter, sinuose, pitted, (42–)64–68(–115) × (4–)9–10(–13)µm. Sexual condition pseudomonoicous;

dwarf male plants on stem rhizoids of female plants;

interior perichaetial leaves abruptly long-acuminate, convolute-sheathing.

straight, erect-spreading, little changed when dry, smooth, 4–6 × 0.2–0.4 mm, most of the leaf tips deciduous and absent, lanceolate, concave proximally, tubulose distally, apex acute;

margins entire or serrulate at apex;

laminae 1-stratose or with 2-stratose regions near apex;

costa long-excurrent, 1/6–1/4 the width of the leaves at base, abaxial ridges absent;

leaf cells smooth, with a row of guide cells, no stereid bands, 1 or 2 rows of cells above and below guide cells with slightly thickened walls and large lumens, the adaxial and abaxial epidermal layers of cells not differentiated;

cell walls between lamina cells not bulging;

alar cells 1-stratose, differentiated, not extending to costa;

proximal laminal cells rectangular to linear, with a few pits, (46–)60–105(–120) × (4–)6–16(–22) µm;

distal laminal cells quadrate, rounded or short-rectangular, not pitted, (12–)17–25(–32) × (8–)10–11(–12) µm. Sexual condition dioicous;

male plants as large as females;

interior perichaetial leaves gradually narrowed to a subulate apex, convolute-sheathing.

Seta

1.5–4 cm, usually aggregate, 3–6 per perichaetium, brown or reddish brown.

1.5–2.5 cm, solitary, yellow to light brown.

Capsule

2–3.5 mm, arcuate, inclined to horizontal, furrowed when dry, yellowish brown or reddish brown;

operculum 2–4 mm.

1.4–2.5 mm, straight and erect, smooth, often irregularly wrinkled when dry, yellow to light brown;

operculum 1–1.8 mm.

Spores

12–24 µm.

10–18 µm.

Dicranum polysetum

Dicranum tauricum

Phenology Capsules mature spring. Capsules mature in summer.
Habitat Commonly on humus, soil over acidic or calcareous rock, and decaying wood in deciduous or more often coniferous forests, occasionally in bogs, fens, and swamps Frequently on rotten logs, stumps, or tree bases in woodlands, sometimes on humus or humus over rock
Elevation 10-2100 m (0-6900 ft) 150-2200 m (500-7200 ft)
Distribution
from FNA
AK; CO; CT; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; PE; QC; SK; YT; Europe; Asia
[WildflowerSearch map]
from FNA
AK; CA; CO; ID; MT; NE; OR; SD; UT; WA; WY; AB; BC; NT; SK; Europe
Discussion

W. L. Peterson (1979) recorded Dicranum polysetum from Nunavut. It is one of the largest, most conspicuous, and most easily recognized species in the genus in North America. The species is immediately recognized by the large stems, 4–15 cm, with a dense mat of whitish or reddish tomentum, giving the stems a thickened appearance, by the shiny, strongly undulate, wide-spreading leaves with margins strongly toothed in the distal half and by the clustered sporophytes (3–6 per perichaetium). Microscopically, the species is readily distinguished by the elongate, abundantly pitted cells throughout the leaf and by the toothed ridges on the abaxial surface of the costa in the distal part of the leaf. Dicranum polysetum is one of the few species in North America that has no apparent intergradations with any other species of the genus. It is common in southeastern Canada, northeastern United States, and the Canadian Rocky Mountains, where it extends south to Wyoming and Colorado, occurring in both states in a small number of localities. It is rare west of the Rockies, where it is known from only a few localities in northwestern Montana, southern Idaho and northeastern Washington and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum tauricum has been reported from Utah by S. Flowers (1973). It is a western North American species, occurring on trees and rotten wood. It is recognized as glossy plants with broken off leaf tips and straight, erect capsules. It can be confused with D. fragilifolium, which also has broken off leaf tips and whose range overlaps with D. tauricum. For distinctions see discussion under 21. D. fragilifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 401. FNA vol. 27, p. 418.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. undulatum, D. viride
Synonyms D. rugosum D. strictum
Name authority Swartz: Monthly Rev., ser. 2, 34: 538. (1801) Sapjegin: Bot. Jahrb. Syst. 46: 10. (1911)
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