Dicranum polysetum |
Dicranum spadiceum |
|
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dicranum moss, wavy broom moss, wavy-leaf heron's-bill moss, wavy-leaf moss |
brown heron's-bill moss, dicranum moss |
|
Habit | Plants in loose tufts, light green, glossy. | Plants in loose to dense tufts, green to yellowish green or brown, glossy. |
Stems | 4–15 cm, densely tomentose with whitish or reddish rhizoids. |
2.5–7 cm, densely tomentose with reddish brown rhizoids. |
Leaves | erect to spreading, ± flexuose, little changed when dry, strongly undulate, (5.5–)7–9.5(–10.5) × 1–2 mm, lanceolate, concave proximally, keeled above, acute; margins strongly toothed in the distal half; laminae 1-stratose; costa ending before apex, occupying ca. 1/16–1/8 of leaf base, strong, with two toothed ridges distally on abaxial surface, with one row of guide cells, two stereid bands, adaxial epidermal layer of cells not differentiated, abaxial layer with a few cells enlarged; cell walls between lamina cells not bulging; leaf cells smooth; alar cells 2-stratose, well-differentiated, not extending to costa; proximal laminal cells elongate, pitted, (45–)78–129(–156) × (5–)9–10(–14) µm; distal laminal cells shorter, sinuose, pitted, (42–)64–68(–115) × (4–)9–10(–13)µm. Sexual condition pseudomonoicous; dwarf male plants on stem rhizoids of female plants; interior perichaetial leaves abruptly long-acuminate, convolute-sheathing. |
straight, erect-spreading, little changed when dry, smooth, (4–)5.5–8(–9) × 0.5–1 mm, concave below, tubulose above, from a lanceolate base to a long-acuminate subula, apices acute or sometimes ± blunt; margins entire or slightly serrate in the distal 1/3; laminae 1-stratose; costa percurrent to shortly excurrent, 1/10–1/6 the width of the leaves at base, smooth to slightly rough above on abaxial surface, abaxial ridges absent, with a row of guide cells, 2 well-developed stereid bands extending almost to apex, abaxial epidermal layer of cells differentiated, rarely some cells in adaxial epidermal layer enlarged; cell walls between lamina cells not or slightly bulging; leaf cells smooth; alar cells 2-stratose, well-differentiated, not extending to costa; proximal laminal cells elongate, pitted, (16–)36–62(–112) × (3–)7–8(–20) µm; distal laminal cells short-rectangular to irregularly angled, pitted, (9–)10–17(–31) × (3)7–8(18) µm. Sexual condition pseudomonoicous; dwarf males on rhizoids of female plants; interior perichaetial leaves abruptly acuminate, convolute-sheathing. |
Seta | 1.5–4 cm, usually aggregate, 3–6 per perichaetium, brown or reddish brown. |
2.5–3.5 cm, solitary, rarely 2 per perichaetium, yellow to reddish yellow. |
Capsule | 2–3.5 mm, arcuate, inclined to horizontal, furrowed when dry, yellowish brown or reddish brown; operculum 2–4 mm. |
2–3.5 mm, arcuate, inclined, furrowed when dry, light to dark brown; operculum 1.5–2.8 mm. |
Spores | 12–24 µm. |
14–22 µm. |
Dicranum polysetum |
Dicranum spadiceum |
|
Phenology | Capsules mature spring. | Capsules mature in summer. |
Habitat | Commonly on humus, soil over acidic or calcareous rock, and decaying wood in deciduous or more often coniferous forests, occasionally in bogs, fens, and swamps | Fens, wet meadows, willow thickets, or humus or soil on or around rocks at lake margins, occasionally drier habitats, such as beach ridges |
Elevation | 10-2100 m (0-6900 ft) | 10-2300 m (0-7500 ft) |
Distribution |
AK; CO; CT; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; PE; QC; SK; YT; Europe; Asia
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AK; CO; MT; UT; WY; AB; BC; MB; NF; NU; ON; QC; SK; YT; Greenland; Europe; Asia |
Discussion | W. L. Peterson (1979) recorded Dicranum polysetum from Nunavut. It is one of the largest, most conspicuous, and most easily recognized species in the genus in North America. The species is immediately recognized by the large stems, 4–15 cm, with a dense mat of whitish or reddish tomentum, giving the stems a thickened appearance, by the shiny, strongly undulate, wide-spreading leaves with margins strongly toothed in the distal half and by the clustered sporophytes (3–6 per perichaetium). Microscopically, the species is readily distinguished by the elongate, abundantly pitted cells throughout the leaf and by the toothed ridges on the abaxial surface of the costa in the distal part of the leaf. Dicranum polysetum is one of the few species in North America that has no apparent intergradations with any other species of the genus. It is common in southeastern Canada, northeastern United States, and the Canadian Rocky Mountains, where it extends south to Wyoming and Colorado, occurring in both states in a small number of localities. It is rare west of the Rockies, where it is known from only a few localities in northwestern Montana, southern Idaho and northeastern Washington and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Dicranum spadiceum has been reported from Saskatchewan by G. Bellolio-Trucco and R. R. Ireland (1990), Colorado by W. L. Peterson (1979), Montana by S. Eversman and A. J. Sharp (1980), and Utah by S. Flowers (1973). It is an arctic-alpine species known principally by the 4–9 mm, straight, erect-spreading leaves, scarcely changed when dry, concave below, tubulose above, narrowed from a lanceolate base to a long-acuminate subula ending in an acute to ± blunt apex; the elongate, pitted cells throughout the leaf; and the costa that is smooth to somewhat rough above on the abaxial surface. Dicranum angustum Lindberg, often considered distinct because of its blunt leaf apices, is considered a synonym after examination of its type from Europe (see discussion by Bellolio-Trucco and Ireland). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 401. | FNA vol. 27, p. 414. |
Parent taxa | Dicranaceae > Dicranum | Dicranaceae > Dicranum |
Sibling taxa | ||
Synonyms | D. rugosum | D. angustum, D. laevidens |
Name authority | Swartz: Monthly Rev., ser. 2, 34: 538. (1801) | J. E. Zetterstedt: Kongl. Svenska Vetensk. Acad. Handl. 5(10): 20. (1865) |
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