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dicranum moss, wavy broom moss, wavy-leaf heron's-bill moss, wavy-leaf moss

dicranum moss, pale-stalk broom-moss, pale-stalk heron's-bill moss

Habit Plants in loose tufts, light green, glossy. Plants in loose tufts, dark green to yellowish green, dull to glossy.
Stems

4–15 cm, densely tomentose with whitish or reddish rhizoids.

1–6 cm, tomentose with white or reddish brown rhizoids.

Leaves

erect to spreading, ± flexuose, little changed when dry, strongly undulate, (5.5–)7–9.5(–10.5) × 1–2 mm, lanceolate, concave proximally, keeled above, acute;

margins strongly toothed in the distal half;

laminae 1-stratose;

costa ending before apex, occupying ca. 1/16–1/8 of leaf base, strong, with two toothed ridges distally on abaxial surface, with one row of guide cells, two stereid bands, adaxial epidermal layer of cells not differentiated, abaxial layer with a few cells enlarged;

cell walls between lamina cells not bulging;

leaf cells smooth;

alar cells 2-stratose, well-differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (45–)78–129(–156) × (5–)9–10(–14) µm;

distal laminal cells shorter, sinuose, pitted, (42–)64–68(–115) × (4–)9–10(–13)µm. Sexual condition pseudomonoicous;

dwarf male plants on stem rhizoids of female plants;

interior perichaetial leaves abruptly long-acuminate, convolute-sheathing.

often secund, straight or nearly so, often crisped when dry, smooth, 4–8 × 0.5–1 mm, lanceolate, acute, tubulose to apex;

margins entire below, serrulate ot serrate near apex;

laminae 1-stratose or sometimes 2-stratose on distal margins;

costa excurrent, 1/5–1/3 the width of the leaves at base, abaxially papillose or toothed from leaf middle to apex, abaxial ridges absent, with a row of guide cells, stereid bands often present and well-developed in proximal part of leaf, absent in distal 1/4–1/3 of leaf, cells above and below guide cells large and thin-walled;

cell walls between lamina cells not or slightly bulging;

leaf cells smooth or often strongly papillose in distal half of leaf;

alar cells 2-stratose, differentiated, often extending to costa, abaxial ridges absent;

proximal laminal cells linear to rectangular, with or without pits, (26–)58–95(–130) × (10–)12–14(–16) µm;

distal laminal cells quadrate, rounded, irregularly angled or short-rectangular, not pitted, (10–)15–26(–38) × (8–)9–11(–12) µm. Sexual condition dioicous;

male plants as large as female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

Seta

1.5–4 cm, usually aggregate, 3–6 per perichaetium, brown or reddish brown.

1–2.5 cm, solitary or sometimes 2, rarely 3, per perichaetium, yellow, rarely reddish yellow or brown with age.

Capsule

2–3.5 mm, arcuate, inclined to horizontal, furrowed when dry, yellowish brown or reddish brown;

operculum 2–4 mm.

1.5–3.5 mm, arcuate, inclined, often with a small struma, strongly furrowed when dry, contracted below mouth, yellow or yellowish brown, rarely reddish yellow or brown with age;

operculum 1.5–2.5 mm.

Spores

12–24 µm.

14–20 µm.

Dicranum polysetum

Dicranum pallidisetum

Phenology Capsules mature spring. Capsules mature in summer.
Habitat Commonly on humus, soil over acidic or calcareous rock, and decaying wood in deciduous or more often coniferous forests, occasionally in bogs, fens, and swamps Humus or soil over rock, rarely rotting logs or decayed wood
Elevation 10-2100 m (0-6900 ft) 500-2000 m (1600-6600 ft)
Distribution
from FNA
AK; CO; CT; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; PE; QC; SK; YT; Europe; Asia
[WildflowerSearch map]
from FNA
AK; ID; MT; OR; WA; AB; BC
Discussion

W. L. Peterson (1979) recorded Dicranum polysetum from Nunavut. It is one of the largest, most conspicuous, and most easily recognized species in the genus in North America. The species is immediately recognized by the large stems, 4–15 cm, with a dense mat of whitish or reddish tomentum, giving the stems a thickened appearance, by the shiny, strongly undulate, wide-spreading leaves with margins strongly toothed in the distal half and by the clustered sporophytes (3–6 per perichaetium). Microscopically, the species is readily distinguished by the elongate, abundantly pitted cells throughout the leaf and by the toothed ridges on the abaxial surface of the costa in the distal part of the leaf. Dicranum polysetum is one of the few species in North America that has no apparent intergradations with any other species of the genus. It is common in southeastern Canada, northeastern United States, and the Canadian Rocky Mountains, where it extends south to Wyoming and Colorado, occurring in both states in a small number of localities. It is rare west of the Rockies, where it is known from only a few localities in northwestern Montana, southern Idaho and northeastern Washington and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum pallidisetum has often been confused with D. fuscescens, being considered a form of it for many years (R. R. Ireland 1966). Unlike that species, which often occurs at low altitudes below 800 m, D. pallidisetum occurs predominately at high elevations, 800–2000 m, rarely being found as low as 500 m. It is distinguished from D. fuscescens by its tubulose leaves, usually 1-stratose or rarely 2-stratose on the margins, by its costa cross section that shows no stereid bands in the distal 1/3 of the leaf, and by its yellow or yellow-brown capsules that are sometimes 2–3 per perichaetium. In contrast, D. fuscescens has leaves keeled above, usually 2-stratose on both margins, rarely 1-stratose on one margin, stereid bands present throughout the leaf and dark brown to reddish brown capsules that are solitary or rarely 2 per perichaetium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 401. FNA vol. 27, p. 413.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
Synonyms D. rugosum D. fuscescens
Name authority Swartz: Monthly Rev., ser. 2, 34: 538. (1801) (J. W. Bailey) Ireland: Bryologist 68: 446. (1965)
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