Dicranum polysetum |
Dicranum majus |
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dicranum moss, wavy broom moss, wavy-leaf heron's-bill moss, wavy-leaf moss |
dicranum moss |
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Habit | Plants in loose tufts, light green, glossy. | Plants in loose tufts, green to light green, glossy to somewhat dull. | ||||
Stems | 4–15 cm, densely tomentose with whitish or reddish rhizoids. |
3–16 cm, naked or with a few whitish rhizoids, rarely moderately tomentose, rhizoids (micronemata) in rows above each leaf. |
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Leaves | erect to spreading, ± flexuose, little changed when dry, strongly undulate, (5.5–)7–9.5(–10.5) × 1–2 mm, lanceolate, concave proximally, keeled above, acute; margins strongly toothed in the distal half; laminae 1-stratose; costa ending before apex, occupying ca. 1/16–1/8 of leaf base, strong, with two toothed ridges distally on abaxial surface, with one row of guide cells, two stereid bands, adaxial epidermal layer of cells not differentiated, abaxial layer with a few cells enlarged; cell walls between lamina cells not bulging; leaf cells smooth; alar cells 2-stratose, well-differentiated, not extending to costa; proximal laminal cells elongate, pitted, (45–)78–129(–156) × (5–)9–10(–14) µm; distal laminal cells shorter, sinuose, pitted, (42–)64–68(–115) × (4–)9–10(–13)µm. Sexual condition pseudomonoicous; dwarf male plants on stem rhizoids of female plants; interior perichaetial leaves abruptly long-acuminate, convolute-sheathing. |
somewhat sparse, falcate-secund or erect-patent, flexuose or straight, little changed when dry, usually smooth, (6–)8–11.5(–15) × 1–2 mm, concave proximally, tubulose above, from a lanceolate to ovate-lanceolate base, gradually narrowed to a long, falcate-secund or straight, acute apex; margins serrate in the distal half, sometimes slightly serrulate above to almost entire; laminae 1-stratose or with few 2-stratose regions on or near the margins; costa percurrent to shortly excurrent, 1/12–1/7 the width of the leaves at base, toothed distally or serrulate to nearly smooth on abaxial surface, with a double row of guide cells that is sometimes interrupted, two stereid bands extending to apex, adaxial epidermal layer of cells with some cells differentiated, the abaxial layer completely differentiated; cell walls between lamina cells not bulging; leaf cells smooth or abaxially prorate or toothed above; alar cells 2-stratose or multistratose, well-differentiated, not extending to costa; proximal laminal cells elongate, pitted, (42–)71–112(–140) × (5–)9–10(–15) µm; distal laminal cells shorter, linear to oval, pitted, (42–)47–61(–99) × (7–)10–11(–17) µm. Sexual condition pseudomonoicous; dwarf males among rhizoids of female plants; interior perichaetial leaves abruptly long-acuminate, convolute-sheathing. |
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Seta | 1.5–4 cm, usually aggregate, 3–6 per perichaetium, brown or reddish brown. |
2.5–5 cm, aggregate, 2–5 per perichaetium, rarely solitary, yellow to light brown. |
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Capsule | 2–3.5 mm, arcuate, inclined to horizontal, furrowed when dry, yellowish brown or reddish brown; operculum 2–4 mm. |
2–3.5 mm, arcuate, inclined to horizontal, smooth to faintly striate when dry, dark brown or yellowish brown; operculum 2–3 mm. |
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Spores | 12–24 µm. |
14–19 µm. |
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Dicranum polysetum |
Dicranum majus |
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Phenology | Capsules mature spring. | |||||
Habitat | Commonly on humus, soil over acidic or calcareous rock, and decaying wood in deciduous or more often coniferous forests, occasionally in bogs, fens, and swamps | |||||
Elevation | 10-2100 m (0-6900 ft) | |||||
Distribution |
AK; CO; CT; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NY; OH; OR; PA; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; PE; QC; SK; YT; Europe; Asia
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nw North America; ne North America; Europe; Asia |
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Discussion | W. L. Peterson (1979) recorded Dicranum polysetum from Nunavut. It is one of the largest, most conspicuous, and most easily recognized species in the genus in North America. The species is immediately recognized by the large stems, 4–15 cm, with a dense mat of whitish or reddish tomentum, giving the stems a thickened appearance, by the shiny, strongly undulate, wide-spreading leaves with margins strongly toothed in the distal half and by the clustered sporophytes (3–6 per perichaetium). Microscopically, the species is readily distinguished by the elongate, abundantly pitted cells throughout the leaf and by the toothed ridges on the abaxial surface of the costa in the distal part of the leaf. Dicranum polysetum is one of the few species in North America that has no apparent intergradations with any other species of the genus. It is common in southeastern Canada, northeastern United States, and the Canadian Rocky Mountains, where it extends south to Wyoming and Colorado, occurring in both states in a small number of localities. It is rare west of the Rockies, where it is known from only a few localities in northwestern Montana, southern Idaho and northeastern Washington and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties ca. 12 (2 in the flora). Dicranum majus is the only North American species of the genus with a double row of guide cells. It is a distinctive species with oceanic tendencies and in North America it commonly grows in northern coastal localities on both sides of the continent, occasionally occurring inland in very moist habitats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 401. | FNA vol. 27, p. 405. | ||||
Parent taxa | Dicranaceae > Dicranum | Dicranaceae > Dicranum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | D. rugosum | |||||
Name authority | Swartz: Monthly Rev., ser. 2, 34: 538. (1801) | Turner: Muscol. Hibern. Spic., 59. (1804) | ||||
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