Dicranum fuscescens |
Dicranum elongatum |
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curly heron's-bill moss, dicranum moss |
dense heron's-bill moss, elongate dicranum moss |
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Habit | Plants light green to dark brownish green, dull, in loose tufts. | Plants in compact tufts, yellowish green to light green, glossy. | ||||
Stems | 1–6(–18) cm, tomen-tose to scarcely tomentose with white or reddish brown rhizoids. |
2–10 cm, tomentose throughout with reddish brown rhizoids. |
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Leaves | falcate-secund, often strongly so, sometimes somewhat sparse, distal leaves curled, proximal leaves with flexuose apices, slightly to strongly curled and crisped when dry, smooth, (3–)4–7(–10) × 0.6–1 mm, concave below, keeled above, from a lanceolate base to a gradually narrowed, fine, keeled subula; margins entire proximally, serrulate to strongly serrate in the distal half, sometimes nearly entire throughout; laminae 1-stratose and usually 2-stratose above on one or both margins, rarely some 2-stratose regions near costa; costa excurrent, 1/6–1/4 the width of the leaves at base, strong, terete, papillose to spinose distally on abaxial surface, occasionally almost smooth, sometimes extending to costa, abaxial ridges absent, with a row of guide cells, 2 well-developed stereid bands extending to distal part of leaf, adaxial epidermal layer of cells not differentiated, abaxial epidermal layer differentiated; cell walls between lamina cells slightly bulging; leaf cells smooth to slightly papillose below on abaxial surface, papillose to spinose prorate above on abaxial surface; alar cells 2-stratose, strongly differentiated, sometimes extending to costa; proximal laminal cells elongate, pitted, (19–)43–62(–93) × (2–)6–8(–12) µm; distal laminal cells irregularly rounded, elliptic, short-rectangular to quadrate, not pitted, (7–)9–23(–36) × (4–)8–12(–14) µm. Sexual condition dioicous; male plants as large as females, growing intermixed, or in separate patches; interior perichaetial leaves abruptly short-acuminate, convolute-sheathing. |
erect-spreading, erect-appressed to rarely spreading when dry, apex sometimes flexuose, smooth, (2.5–)3–4.5(–6) × 0.3–0.5 mm, from a lanceolate base to a long, fine, tubulose subula, apex usually acute, sometimes apical leaves of stems blunt, julaceous; margins entire, rarely slightly denticulate at apex; laminae 1-stratose; costa percurrent or shortly excurrent, 1/6–1/4 the width of the leaves at base, smooth or weakly rough above on abaxial surface, abaxial ridges absent, with a row of guide cells, two stereid bands not extending to apex, adaxial and abaxial epidermal layers of cells usually not differentiated or sometimes a few cells in both layers enlarged; cell walls between lamina cells slightly bulging; leaf cells smooth; alar cells 1-stratose or 2-stratose, well-differentiated, not extending to costa; proximal laminal cells usually elongate, pitted, (14–)24–36(–49) × (2–)4–6(–9) µm; distal laminal cells short-rectangular, quadrate, rounded, or sometimes irregularly shaped, incrassate, not pitted, (5–)12–17(–23) × (4–)6–8(–9) µm. Sexual condition dioicous; male plants as large as females; interior perichaetial leaves abruptly short-acuminate, convolute-sheathing. |
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Seta | 1–3.5 cm, solitary, yellow, reddish yellow or brown. |
1.5–2 cm., solitary, yellowish to yellowish brown or reddish yellow. |
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Capsule | 1–3 mm, arcuate, inclined to horizontal, strumose, strongly furrowed when dry, somewhat contracted below mouth, dark brown to reddish brown; operculum 1–2 mm. |
1.2–1.8 mm, nearly straight and erect to slightly arcuate, striate when dry, yellowish brown; operculum 1.2–1.8 mm. |
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Spores | 14–24 µm. |
17–22 µm. |
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Dicranum fuscescens |
Dicranum elongatum |
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Phenology | Capsules mature in summer. | |||||
Habitat | Common in arctic or alpine tundra, soil, rocks or soil over rocks, rock ledges, cliff shelves, rarely decayed wood and stumps, sometimes in bogs and fens | |||||
Elevation | 30-3700 m (100-12100 ft) | |||||
Distribution | North America; Europe; Asia
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AK; CO; ME; MN; MT; NH; NY; AB; BC; MB; NF; NS; NT; NU; ON; QC; SK; YT; Greenland; Europe; Asia |
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Discussion | Varieties ca. 7 (2 in the flora). l. Plants small, stems 1–6(–10) cm, usually densely foliate; leaf margins serrulate to strongly serrate in distal half; costa papillose to spinose distally on abaxial surface; distal laminal cells short-rectangular to quadrate, mostly 18–23 µm long; seta mostly shorter than 2.5 cm 15a. Dicranum fuscescens var. fuscescens (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Dicranum elongatum is easy to recognize because the plants are relatively small, slender, glossy, and grow in dense, compact tufts. The leaves are short, narrow, usually less than 6 × 0.5 mm, ending in a long, fine, tubulose subula, acute at the apex, smooth or nearly so on the abaxial surface, and have entire margins or rarely some denticulations at the apex. Capsules, which are not uncommon on plants in some arctic localities, are short, less than 2 mm, straight and erect to slightly arcuate, smooth but striate when dry. Dicranum elongatum can readily be confused with the closely related D. groenlandicum. They both are arctic-alpine species sharing nearly the same type of habitat with almost the same distribution. They can usually be readily distinguished in their typical form but there appear to be many intergradations that cause difficulty. Dicranum groenlandicum has narrowly obtuse leaf apices and cells that are pitted to well above the middle of the leaf in contrast to the acute leaf apices and cells that are pitted only up to the middle of the leaf in D. elongatum. E. Hegewald (1972) found that the pits in the leaf are the most reliable means of separating difficult specimens of Finnish plants. H. A. Crum and L. E. Anderson (1981) expressed doubt as to whether the two are genetically different. Both D. elongatum and D. groenlandicum often bear at the apex of some stems julaceous shoots with small, ovate, broad leaves with obtuse apices. This can lead to the misidentification of some plants of D. elongatum unless the leaves in the basal part of the stems are always used to determine the shape of the leaf apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 411. | FNA vol. 27, p. 415. | ||||
Parent taxa | Dicranaceae > Dicranum | Dicranaceae > Dicranum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Turner: Muscol. Hibern. Spic., 60, plate 5, fig. 1. (1804) | Schwägrichen: Sp. Musc. Frond. Suppl. 1(1): 171, plate 43. (1811) | ||||
Web links |