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condensed dicranum moss

undulate dicranum moss, wavy heron's-bill moss

Habit Plants in ± dense tufts, light green to yellowish brown, dull. Plants in dense compact tufts, green to yellowish brown, shiny, rarely somewhat dull.
Stems

1–9 cm, densely tomentose below with reddish brown rhizoids.

3–8(–17) cm, densely tomentose with reddish brown rhizoids.

Leaves

erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex;

margins serrulate in the distal half, involute in central part;

laminae 1-stratose, with a few 2-stratose regions;

costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells strongly bulging;

leaf cells papillose above on abaxial surface, sometimes nearly smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex;

distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

erect-appressed or sometimes slightly falcate or flexuose, somewhat contorted when dry, the apex often twisted, weakly to strongly undulate, (4.5–)5–7.5(–11) × 0.7–1.2 mm, concave proximally, keeled above, lanceolate to oblong-lanceolate, broadly acute, rarely narrowly acute;

margins plane, sometimes involute at base, broadly recurved at apex, entire proximally, serrulate to serrate in the distal half;

laminae 1-stratose, sometimes with a few 2-stratose regions;

costa strong, ending before the apex, sometimes nearly percurrent, 1/6–1/3 the width of the leaves at base, smooth to serrulate above on abaxial surface, with a row of guide cells, two stereid bands extending to apex, adaxial epidermal layer of cells not differentiated, the abaxial layer diffferentiated;

cell walls between lamina cells weakly to strongly bulging abaxially and adaxially;

leaf cells smooth to ± papillose near apex on abaxial surface;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, incrassate, pitted, (22–)38–61(–79) × (2–)4–6(–8) µm;

median laminal cells rectangular, pitted;

distal laminal cells short-rectangular to irregularly angled, not pitted, (7–)11–14(–28) × (4–)11–12(–17) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly acuminate, convolute-sheathing.

Seta

1–4 cm, solitary, rarely 2 per perichaetium, yellowish.

2–4 cm, solitary, yellow to brown.

Capsule

1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown;

operculum 1.5–2.5 mm.

2–2.8 mm, arcuate, inclined, furrowed when dry, ± contracted below mouth, yellow to yellowish brown or reddish brown;

operculum 2–3 mm.

Spores

19–26 µm.

14–28 µm.

Dicranum condensatum

Dicranum undulatum

Phenology Capsules mature spring. Capsules mature spring.
Habitat Exposed sandy soil, often in pine woods, frequently over sandstone and limestone Usually in wet habitats, especially bogs in Sphagnum hummocks, fens, swamps, marshes, margins of lakes, sometimes on or among rocks, rock outcrops and cliffs, occasionally in mesic woods or on dry bluffs on soil, rotten wood, or humus
Elevation 0-700 m (0-2300 ft) 10-2200 m (0-7200 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
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from FNA
AK; CA; CT; IN; MA; ME; MI; MN; MT; NC; NH; NJ; NY; OH; PA; SD; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Europe; Asia
Discussion

Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

R. R. Ireland (1971b) recorded Dicranum undulatum from Colorado, but W. Weber (pers. comm.) believes this is a misidentification for D. polysetum. Dicranum undulatum is primarily a boreal species that is distinctive because of the yellowish green or yellowish brown, shiny, erect-appressed, lanceolate to oblong-lanceolate, undulate, keeled leaves, the leaf margins that are serrulate to serrate, and broadly recurved in the distal half, the leaf apices that are usually broadly acute, rarely narrowly acute, and the costae that are mostly subpercurrent or sometimes percurrent, smooth to serrulate above on abaxial surface. When the leaf apices are narrow, ± acute, and the costa percurrent, as some D. undulatum plants are on rare occasion, the species when sterile can be confused with D. ontariense. Dicranum undulatum has shiny leaves with twisted apices when dry compared to the more dull leaves that are variously contorted in D. ontariense. When sporophytes are present the aggregate setae (2–5 per perichaetium) of D. ontariense will immediately distinguish it from D. undulatum which has solitary setae. This species recently has been introduced on the University of California Berkeley Campus in a lawn-landscaping area; this is the only California locality where it has been found. The plants are sterile but appear morphologically similar to those growing in native habitats in other parts of North America. Dicranum undulatum of G. F. Weber & C. Mohr (1803) is not a synonym of the taxon treated here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 406.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. viride
Synonyms D. sabuletorum D. bergeri
Name authority Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, Bridel: J. Bot. (Schrader) 1800(2): 294. (1801)
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