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condensed dicranum moss

broken-leaf heron's-bill moss, broken-leaf moss, dicranum moss

Habit Plants in ± dense tufts, light green to yellowish brown, dull. Plants in dense tufts, light green to yellowish green, glossy.
Stems

1–9 cm, densely tomentose below with reddish brown rhizoids.

0.5–3 cm, tomentose below with whitish or reddish brown rhizoids.

Leaves

erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex;

margins serrulate in the distal half, involute in central part;

laminae 1-stratose, with a few 2-stratose regions;

costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells strongly bulging;

leaf cells papillose above on abaxial surface, sometimes nearly smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex;

distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

straight, erect-spreading, little changed when dry, smooth, 4–6 × 0.2–0.4 mm, most of the leaf tips deciduous and absent, lanceolate, concave proximally, tubulose distally, apex acute;

margins entire or serrulate at apex;

laminae 1-stratose or with 2-stratose regions near apex;

costa long-excurrent, 1/6–1/4 the width of the leaves at base, abaxial ridges absent;

leaf cells smooth, with a row of guide cells, no stereid bands, 1 or 2 rows of cells above and below guide cells with slightly thickened walls and large lumens, the adaxial and abaxial epidermal layers of cells not differentiated;

cell walls between lamina cells not bulging;

alar cells 1-stratose, differentiated, not extending to costa;

proximal laminal cells rectangular to linear, with a few pits, (46–)60–105(–120) × (4–)6–16(–22) µm;

distal laminal cells quadrate, rounded or short-rectangular, not pitted, (12–)17–25(–32) × (8–)10–11(–12) µm. Sexual condition dioicous;

male plants as large as females;

interior perichaetial leaves gradually narrowed to a subulate apex, convolute-sheathing.

Seta

1–4 cm, solitary, rarely 2 per perichaetium, yellowish.

1.5–2.5 cm, solitary, yellow to light brown.

Capsule

1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown;

operculum 1.5–2.5 mm.

1.4–2.5 mm, straight and erect, smooth, often irregularly wrinkled when dry, yellow to light brown;

operculum 1–1.8 mm.

Spores

19–26 µm.

10–18 µm.

Dicranum condensatum

Dicranum tauricum

Phenology Capsules mature spring. Capsules mature in summer.
Habitat Exposed sandy soil, often in pine woods, frequently over sandstone and limestone Frequently on rotten logs, stumps, or tree bases in woodlands, sometimes on humus or humus over rock
Elevation 0-700 m (0-2300 ft) 150-2200 m (500-7200 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
[WildflowerSearch map]
from FNA
AK; CA; CO; ID; MT; NE; OR; SD; UT; WA; WY; AB; BC; NT; SK; Europe
Discussion

Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum tauricum has been reported from Utah by S. Flowers (1973). It is a western North American species, occurring on trees and rotten wood. It is recognized as glossy plants with broken off leaf tips and straight, erect capsules. It can be confused with D. fragilifolium, which also has broken off leaf tips and whose range overlaps with D. tauricum. For distinctions see discussion under 21. D. fragilifolium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 418.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. undulatum, D. viride
Synonyms D. sabuletorum D. strictum
Name authority Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, Sapjegin: Bot. Jahrb. Syst. 46: 10. (1911)
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