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condensed dicranum moss

dicranum moss

Habit Plants in ± dense tufts, light green to yellowish brown, dull. Plants in loose tufts, green to yellowish green or yellowish brown, dull.
Stems

1–9 cm, densely tomentose below with reddish brown rhizoids.

2–11 cm, densely tomentose below with reddish brown rhizoids, interruptedly foliate, the leaves crowded in some parts forming swollen, globose regions.

Leaves

erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex;

margins serrulate in the distal half, involute in central part;

laminae 1-stratose, with a few 2-stratose regions;

costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells strongly bulging;

leaf cells papillose above on abaxial surface, sometimes nearly smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex;

distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

erect-spreading, arched and loosely imbricate with contorted apices when dry, ± undulate to rugose, (2.5–)5–6.5(–9) × 1–1.7 mm, concave below, tubulose above, acute, ovate at base, widest point at or somewhat below middle;

margins serrulate to serrate in distal half, plane to involute;

laminae 1-stratose or with 2-stratose regions near costa and on margins;

costa percurrent to shortly excurrent, 1/8–1/4 the width of the leaves at base, serrulate or toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two stereid bands extending to apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells weakly to strongly bulging;

leaf cells strongly papillose above on abaxial surface, sometimes a few papillae on adaxial surface;

alar cells 2-stratose, distinctly differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (30–)49–68(–101) × (5–)10–11(–15) µm, abruptly shorter in distal half of leaf;

distal laminal cells short, not or indistinctly pitted, cell walls irregularly thickened, (7–)11–12(–23) × (5–)7–13(–20) µm. Sexual condition pseudomonoicous;

dwarf males on rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

Seta

1–4 cm, solitary, rarely 2 per perichaetium, yellowish.

1–3 cm, solitary, reddish yellow or brown.

Capsule

1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown;

operculum 1.5–2.5 mm.

1.5–2.5 mm, arcuate, inclined to horizontal, contracted below mouth and furrowed when dry, yellowish brown to brown;

operculum 1.5–2.5 mm.

Spores

19–26 µm.

16–24 µm.

Dicranum condensatum

Dicranum spurium

Phenology Capsules mature spring. Capsules mature spring.
Habitat Exposed sandy soil, often in pine woods, frequently over sandstone and limestone Sandy soil, decayed logs, acidic rock, humus over exposed bluffs or outcrops, in pine woods, sometimes bogs
Elevation 0-700 m (0-2300 ft) 0-1400 m (0-4600 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
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from FNA
AR; CT; GA; IL; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; OK; PA; TN; VA; VT; WV; NF; NS; ON; PE; QC; Europe
[WildflowerSearch map]
Discussion

Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum spurium is easily recognized by the turgid aspect of the plants with arched, imbricate leaves and with interruptedly foliate stems, i.e., some regions that appear swollen or globose as a result of several growth periods. It is likely to be confused only with 11. D. condensatum (see discussion thereunder).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 409.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. tauricum, D. undulatum, D. viride
Synonyms D. sabuletorum
Name authority Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, Hedwig: Sp. Musc. Frond., 141. (1801)
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