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condensed dicranum moss

dicranum moss

Habit Plants in ± dense tufts, light green to yellowish brown, dull. Plants in dense tufts, green to yellowish green or brownish, ± glossy.
Stems

1–9 cm, densely tomentose below with reddish brown rhizoids.

2–8 cm, tomentose with reddish brown rhizoids.

Leaves

erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex;

margins serrulate in the distal half, involute in central part;

laminae 1-stratose, with a few 2-stratose regions;

costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells strongly bulging;

leaf cells papillose above on abaxial surface, sometimes nearly smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex;

distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

straight or nearly so, spreading, little changed when dry, smooth, 3–5.5 × 0.6–1.2 mm, concave or tubulose proximally, tubulose to slightly keeled above; ovate-lanceolate, apex obtusely acute;

margins serrate near apex;

laminae 1-stratose;

costa subpercurrent to percurrent, 1/10–1/8 the width of the leaves at base, smooth or weakly toothed on abaxial surface near apex, with a row of guide cells, two weak stereid bands, at least in basal part of leaf, adaxial and abaxial epidermal layers of cells not differentiated;

cell walls between lamina cells not bulging;

leaf cells smooth;

alar cells 1- or 2-stratose in part, differentiated, not extending to costa;

proximal laminal cells linear-rectangular, pitted, (45–)65–120(–150) × (13–)16–17(–19) µm;

distal laminal cells shorter, narrow, pitted or with few pits, (20–)30–45(–60) × (5–)8–10(–13) µm. Sexual condition dioicous;

male plants about as large as the female or slightly smaller;

interior perichaetial leaves ± abruptly acuminate, convolute-sheathing.

Seta

1–4 cm, solitary, rarely 2 per perichaetium, yellowish.

1.5–3 cm, solitary, rarely two per perichaetium, yellow to reddish brown.

Capsule

1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown;

operculum 1.5–2.5 mm.

1.5–4 mm, erect, straight or nearly so, furrowed when dry, brown;

operculum 1.6–2.8 mm.

Spores

19–26 µm.

13–19 µm.

Dicranum condensatum

Dicranum rhabdocarpum

Phenology Capsules mature spring. Capsules mature summer.
Habitat Exposed sandy soil, often in pine woods, frequently over sandstone and limestone Soil, soil over rock, peaty soil or rotten wood
Elevation 0-700 m (0-2300 ft) 1600-3300 m (5200-10800 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
[WildflowerSearch map]
from FNA
AZ; CO; NM; WY; Mexico (Chihuahua, Tamaulipas); West Indies (Dominican Republic); Central America (Guatemala)
Discussion

Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum rhabdocarpum is an easily recognized species that occurs in the flora area only in the Rocky Mountains and the mountains of Arizona. It is the only species in the section Dicranum that has erect, straight to weakly arcuate capsules. Other important distinguishing features are the ovate-lanceolate, straight, obtusely acute, 1-stratose leaves with alar cells 1- or 2-stratose in part and the subpercurrent to percurrent costae that are smooth or weakly toothed on the abaxial surface near the leaf apex. Dicranum rhabdocarpum has been placed in the segregate genus Orthodicranum by J. M. Holzinger (1925b) and other bryologists mainly because of its straight and erect capsules. However, it differs from taxa commonly placed in that genus, i.e., D. flagellare, D. fulvum, D. montanum, D. strictum, and D. viride, by the elongate, pitted cells and the alar cells that are sometimes 2-stratose.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 406.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
Synonyms D. sabuletorum Orthodicranum rhabdocarpum
Name authority Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, Sullivant: Mem. Amer. Acad. Arts, n. s. 4: 172. (1849)
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