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condensed dicranum moss

dicranum moss

Habit Plants in ± dense tufts, light green to yellowish brown, dull. Plants in loose tufts, yellowish green, glossy.
Stems

1–9 cm, densely tomentose below with reddish brown rhizoids.

5–8 cm, scarcely tomentose with whitish to reddish brown rhizoids.

Leaves

erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex;

margins serrulate in the distal half, involute in central part;

laminae 1-stratose, with a few 2-stratose regions;

costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells strongly bulging;

leaf cells papillose above on abaxial surface, sometimes nearly smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex;

distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

erect or spreading, distal leaves with twisted apices when dry, slightly rugose, (3.5–)5–7(–8) × 1–1.5 mm, concave proximally, tubulose distally, from an ovate-lanceolate base to a long or short subula, broadly acute, some leaves short, ovate, somewhat obtuse, forming julaceous regions, in the middle or basal part of the stems;

margins not or slightly serrate in the distal part, somewhat involute in the middle part;

laminae 1-stratose;

costa ending before the apex, 1/13–1/8 the width of the leaves at base, smooth or sometimes the distal part with two weakly serrated ridges on abaxial surface, row of guide cells, two stereid bands, adaxial and abaxial epidermal layers of cells not differentiated or sometimes a few cells in the abaxial layer differentiated;

cell walls between lamina cells not bulging;

leaf cells smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate-sinuose, pitted, (42–)75–87(–107) × (7–)8–10(–16) µm;

distal laminal cells short, sinuose, pitted, (16–)30–65(–79) × (7–)9–13(–16) µm. Sexual condition pseudomonoicous;

dwarf males on rhizoids of female plants;

interior perichaetial leaves abruptly long-acuminate, convolute-sheathing.

Seta

1–4 cm, solitary, rarely 2 per perichaetium, yellowish.

2–4 cm, solitary, yellow, yellowish brown or reddish yellow.

Capsule

1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown;

operculum 1.5–2.5 mm.

2–3.5 mm, arcuate, inclined, smooth, ± furrowed when dry, yellow to yellowish brown;

operculum 1.5–2 mm.

Spores

19–26 µm.

14–24 µm.

Dicranum condensatum

Dicranum leioneuron

Phenology Capsules mature spring. Capsules mature spring.
Habitat Exposed sandy soil, often in pine woods, frequently over sandstone and limestone Primarily in hummocks in ombrotrophic and oligotrophic peatlands
Elevation 0-700 m (0-2300 ft) 10-1100 m (0-3600 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
[WildflowerSearch map]
from FNA
AK; BC; NB; NF; NS; NT; ON; PE; QC; YT; nw Europe
Discussion

Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum leioneuron is apparently closely related to D. bonjeanii. It is best distinguished by the subulate leaves with twisted apices when dry, the usual occurrence of julaceous stem portions with broad, short, concave, and somewhat obtuse leaves in contrast to the longer, narrower, subulate, and acute leaves of the other stem portions. Its occurrence only in hummocks in peatlands also is a clue to its identity. The plants rarely produce sporophytes but in any event they cannot be distinguished from those of the other species in section Dicranum. The species probably occurs in peatlands in northeastern United States and it should be looked for in that region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 404.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
Synonyms D. sabuletorum
Name authority Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, Kindberg: Bull. Torrey Bot. Club 16: 92. (1889)
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