Dicranum condensatum |
Dicranum fuscescens |
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condensed dicranum moss |
curly heron's-bill moss, dicranum moss |
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Habit | Plants in ± dense tufts, light green to yellowish brown, dull. | Plants light green to dark brownish green, dull, in loose tufts. | ||||
Stems | 1–9 cm, densely tomentose below with reddish brown rhizoids. |
1–6(–18) cm, tomen-tose to scarcely tomentose with white or reddish brown rhizoids. |
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Leaves | erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex; margins serrulate in the distal half, involute in central part; laminae 1-stratose, with a few 2-stratose regions; costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated; cell walls between lamina cells strongly bulging; leaf cells papillose above on abaxial surface, sometimes nearly smooth; alar cells 2-stratose, differentiated, not extending to costa; proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex; distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous; dwarf males on stem rhizoids of female plants; interior perichaetial leaves abruptly short-acuminate, convolute-sheathing. |
falcate-secund, often strongly so, sometimes somewhat sparse, distal leaves curled, proximal leaves with flexuose apices, slightly to strongly curled and crisped when dry, smooth, (3–)4–7(–10) × 0.6–1 mm, concave below, keeled above, from a lanceolate base to a gradually narrowed, fine, keeled subula; margins entire proximally, serrulate to strongly serrate in the distal half, sometimes nearly entire throughout; laminae 1-stratose and usually 2-stratose above on one or both margins, rarely some 2-stratose regions near costa; costa excurrent, 1/6–1/4 the width of the leaves at base, strong, terete, papillose to spinose distally on abaxial surface, occasionally almost smooth, sometimes extending to costa, abaxial ridges absent, with a row of guide cells, 2 well-developed stereid bands extending to distal part of leaf, adaxial epidermal layer of cells not differentiated, abaxial epidermal layer differentiated; cell walls between lamina cells slightly bulging; leaf cells smooth to slightly papillose below on abaxial surface, papillose to spinose prorate above on abaxial surface; alar cells 2-stratose, strongly differentiated, sometimes extending to costa; proximal laminal cells elongate, pitted, (19–)43–62(–93) × (2–)6–8(–12) µm; distal laminal cells irregularly rounded, elliptic, short-rectangular to quadrate, not pitted, (7–)9–23(–36) × (4–)8–12(–14) µm. Sexual condition dioicous; male plants as large as females, growing intermixed, or in separate patches; interior perichaetial leaves abruptly short-acuminate, convolute-sheathing. |
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Seta | 1–4 cm, solitary, rarely 2 per perichaetium, yellowish. |
1–3.5 cm, solitary, yellow, reddish yellow or brown. |
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Capsule | 1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown; operculum 1.5–2.5 mm. |
1–3 mm, arcuate, inclined to horizontal, strumose, strongly furrowed when dry, somewhat contracted below mouth, dark brown to reddish brown; operculum 1–2 mm. |
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Spores | 19–26 µm. |
14–24 µm. |
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Dicranum condensatum |
Dicranum fuscescens |
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Phenology | Capsules mature spring. | |||||
Habitat | Exposed sandy soil, often in pine woods, frequently over sandstone and limestone | |||||
Elevation | 0-700 m (0-2300 ft) | |||||
Distribution |
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
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North America; Europe; Asia
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Discussion | Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties ca. 7 (2 in the flora). l. Plants small, stems 1–6(–10) cm, usually densely foliate; leaf margins serrulate to strongly serrate in distal half; costa papillose to spinose distally on abaxial surface; distal laminal cells short-rectangular to quadrate, mostly 18–23 µm long; seta mostly shorter than 2.5 cm 15a. Dicranum fuscescens var. fuscescens (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27, p. 409. | FNA vol. 27, p. 411. | ||||
Parent taxa | Dicranaceae > Dicranum | Dicranaceae > Dicranum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | D. sabuletorum | |||||
Name authority | Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, | Turner: Muscol. Hibern. Spic., 60, plate 5, fig. 1. (1804) | ||||
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