The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

condensed dicranum moss

fragile leaf dicranum moss

Habit Plants in ± dense tufts, light green to yellowish brown, dull. Plants in compact tufts, light green to yellowish brown, glossy.
Stems

1–9 cm, densely tomentose below with reddish brown rhizoids.

1.5–6 cm, densely tomentose with dark brown to red rhizoids.

Leaves

erect-spreading, flexuose, ± curled at the tips when dry, slightly undulate or rugose, (2–)3.5–4.5(–7) × 0.5–1 mm, concave below, strongly keeled above, oblong-lanceolate, with a broad lanceolate base gradually narrowing to a short-acuminate apex;

margins serrulate in the distal half, involute in central part;

laminae 1-stratose, with a few 2-stratose regions;

costa percurrent to shortly excurrent, 1/10–1/5 the width of the leaves at base, strong, smooth or papillose to toothed above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thick stereid bands extending well into the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells strongly bulging;

leaf cells papillose above on abaxial surface, sometimes nearly smooth;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, pitted, (25–)41–56(–112) × (5–)6–7(–10) µm, abruptly shorter towards the apex;

distal laminal cells quadrate, irregularly angled, rounded or rectangular, not or indistinctly pitted, cell walls unequally thickened, (5–)7–8(–20) × (5–)7–8(–12) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

straight, erect-spreading, rigid, appressed when dry, smooth, (5–)6–7(–7.5) × 0.4–0.6 mm, most of the leaf tips deciduous and absent, concave proximally, canaliculate distally, from a lanceolate base to a long subula formed by the excurrent costa, apex acute;

margins entire to somewhat serrulate above;

laminae 1-stratose or some 2-stratose regions near costa;

costa excurrent, 1/4–1/3 the width of the leaves at base, smooth or slightly rough above on abaxial surface, abaxial ridges absent, with a row of guide cells, two thin stereid bands (2–3 cells thick), adaxial and abaxial epidermal layers not differentiated or sometimes with a few cells in both layers enlarged;

cell walls between lamina cells slightly bulging;

leaf cells smooth;

alar cells 1-stratose, sometimes with some 2-stratose regions, well-differentiated, sometimes extending to costa;

proximal laminal cells elongate-rectangular, usually pitted or indistinctly pitted, (25–)39–55(–84) × (5–)7–8(–10) µm;

median laminal cells rectangular, not pitted, (11–)21–22(–37) × (4–)7–8(–10) µm;

distal laminal cells nearly elliptic, incrassate.

Seta

1–4 cm, solitary, rarely 2 per perichaetium, yellowish.

1.5–2.5 cm, solitary, yellowish to brown.

Sexual condition

dioicous;

male plants as tall as females, usually more slender;

interior perichaetial leaves abruptly long-acuminate, convolute-sheathing.

Capsule

1.5–3 mm, arcuate, inclined to horizontal, furrowed when dry, slightly contracted below the mouth, yellowish brown;

operculum 1.5–2.5 mm.

1.8–2 mm, arcuate to nearly straight, ± erect, smooth, sometimes striate when dry, yellowish brown;

operculum 1–2 mm.

Spores

19–26 µm.

16–28 µm.

Dicranum condensatum

Dicranum fragilifolium

Phenology Capsules mature spring. Capsules mature in summer.
Habitat Exposed sandy soil, often in pine woods, frequently over sandstone and limestone Rotten logs and stumps, humic soil, humus over rocks, usually in coniferous woods, occasionally in mixed woods and bogs
Elevation 0-700 m (0-2300 ft) 30-1900 m (100-6200 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; NB; NS; QC
[WildflowerSearch map]
from FNA
AK; MI; AB; BC; MB; NF; NT; ON; QC; SK; YT; Greenland; Europe; Asia
Discussion

Dicranum condensatum is a common species recognized in the field by its occurrence on sandy soil, by its light green to yellowish brown, dull plants in compact, dense tufts, its erect-spreading, oblong-lanceolate leaves, narrowing to a short-acuminate apex, keeled above, somewhat undulate or rugose, curled at tips when dry, and its solitary setae with an arcuate, inclined to horizontal capsule. The papillose leaf cells with strongly bulging walls between the cells and the irregularly angled median and distal cells with nonpitted walls are distinctive microscopic characters. Dicranum spurium is the species most commonly confused with D. condensatum. Both species occur on sandy soil with their ranges overlapping. The former occurs farther northward, where it is found in Newfoundland, while the latter occurs farther to the south, where its range extends to the tip of Florida. Dicranum spurium has broader leaves, the broadest point near the middle rather than near the base like D. condensatum, and leaves that are tubulose above (most noticeable when wet) rather than keeled. Rarely, D. condensatum can be mistaken for 9. D. ontariense as discussed under that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dicranum fragilifolium is a rare boreal species with shiny, light green to yellowish brown, erect-spreading leaves. The distal portion of the leaves is fragile and often broken off, thereby giving the plants a distinctive appearance. The deciduous leaf apices presumably serve as a type of asexual reproduction by regenerating to produce new plants. The species is often confused with D. tauricum, another species with deciduous leaf tips, but the latter occurs only in western North America. Where their ranges overlap they frequently are difficult to tell apart. When sporophytes are present the straight capsules of D. tauricum are distinctive from the usually arcuate ones of D. fragilifolium, which unfortunately rarely produces them. When sterile, the best way to separate the two is by the costa cross section in the proximal half of the leaf: D. fragilifolium has stereid cells in two thin bands while D. tauricum has none. Also, in D. fragilifolium there are 2–3 layers of cells above and below the guide cells, while in D. tauricum there are 1 or rarely 2 layers of cells. One other species with deciduous leaf tips that has sometimes been confused with D. fragilifolium is D. viride of eastern North America. It has a straight capsule like D. tauricum but as in D. fragilifolium capsules are rarely produced. The broad costa of D. viride, covering 1/3 or more of the leaf base, will distinguish it from D. fragilifolium, the costa of which covers 1/4 or less of the leaf base.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 409. FNA vol. 27, p. 416.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. acutifolium, D. bonjeanii, D. brevifolium, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
Synonyms D. sabuletorum
Name authority Hedwig: Sp.. Musc. Frond., 139. 1801, not plate 34, Lindberg: Bot. Not. 1857: 147. (1857)
Web links