The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

acuteleaf dicranum moss, sharp-leaf heron's-bill moss

undulate dicranum moss, wavy heron's-bill moss

Habit Plants in dense to loose tufts, light green to light brown, dull. Plants in dense compact tufts, green to yellowish brown, shiny, rarely somewhat dull.
Stems

1.5–7 cm, generally branched above, somewhat tomentose with reddish brown rhizoids.

3–8(–17) cm, densely tomentose with reddish brown rhizoids.

Leaves

erect-spreading, slightly curled to ± straight when dry, usually with a few undulations in distal part, (3.5–)5–8(–10) mm × 0.5–1 mm, concave below, keeled above, sometimes margins ± involute, lanceolate, narrowly acute to acuminate;

margins serrulate to serrate above;

laminae 1-stratose or with few 2-stratose regions on or near the distal margins;

costa percurrent to shortly excurrent, 1/6–1/4 the width of the leaves at base, smooth to slightly papillose above on abaxial surface, abaxial ridges absent, with a row of guide cells, two stereid bands extending to the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated;

cell walls between lamina cells not or slightly bulging;

leaf cells smooth to weakly papillose above on abaxial surface;

alar cells 2-stratose, differentiated, often extending to costa;

proximal laminal cells linear-rectangular, pitted, (22–)41–60(–90) × (5–)6–8(–12) µm;

distal laminal cells short-rectangular, irregularly angled or rounded, not pitted, (7–)12–20(–32) × (4–)8–9(–14) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly short-acuminate, convolute-sheathing.

erect-appressed or sometimes slightly falcate or flexuose, somewhat contorted when dry, the apex often twisted, weakly to strongly undulate, (4.5–)5–7.5(–11) × 0.7–1.2 mm, concave proximally, keeled above, lanceolate to oblong-lanceolate, broadly acute, rarely narrowly acute;

margins plane, sometimes involute at base, broadly recurved at apex, entire proximally, serrulate to serrate in the distal half;

laminae 1-stratose, sometimes with a few 2-stratose regions;

costa strong, ending before the apex, sometimes nearly percurrent, 1/6–1/3 the width of the leaves at base, smooth to serrulate above on abaxial surface, with a row of guide cells, two stereid bands extending to apex, adaxial epidermal layer of cells not differentiated, the abaxial layer diffferentiated;

cell walls between lamina cells weakly to strongly bulging abaxially and adaxially;

leaf cells smooth to ± papillose near apex on abaxial surface;

alar cells 2-stratose, differentiated, not extending to costa;

proximal laminal cells elongate, incrassate, pitted, (22–)38–61(–79) × (2–)4–6(–8) µm;

median laminal cells rectangular, pitted;

distal laminal cells short-rectangular to irregularly angled, not pitted, (7–)11–14(–28) × (4–)11–12(–17) µm. Sexual condition pseudomonoicous;

dwarf males on stem rhizoids of female plants;

interior perichaetial leaves abruptly acuminate, convolute-sheathing.

Seta

1.2–2.5 cm, solitary, yellow to reddish yellow.

2–4 cm, solitary, yellow to brown.

Capsule

2–2.7 mm, arcuate, inclined to horizontal, furrowed when dry, rarely slightly strumose, brown to reddish brown;

operculum 1.5–2.5 mm.

2–2.8 mm, arcuate, inclined, furrowed when dry, ± contracted below mouth, yellow to yellowish brown or reddish brown;

operculum 2–3 mm.

Spores

14–28 µm.

14–28 µm.

Dicranum acutifolium

Dicranum undulatum

Phenology Capsules mature summer. Capsules mature spring.
Habitat Somewhat calcareous soil, boulders, rock outcrops and cliff ledges, sometimes humus, rarely rotten logs Usually in wet habitats, especially bogs in Sphagnum hummocks, fens, swamps, marshes, margins of lakes, sometimes on or among rocks, rock outcrops and cliffs, occasionally in mesic woods or on dry bluffs on soil, rotten wood, or humus
Elevation 700-2500 m (2300-8200 ft) 10-2200 m (0-7200 ft)
Distribution
from FNA
AK; ME; MT; NH; NM; NY; WY; AB; BC; MB; NF; NT; NU; ON; QC; YT; Greenland; Europe; Asia
from FNA
AK; CA; CT; IN; MA; ME; MI; MN; MT; NC; NH; NJ; NY; OH; PA; SD; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT; Greenland; Europe; Asia
Discussion

Dicranum acutifolium has been reported from Colorado by W. A. Weber (1973), but has been deleted from the flora (Weber, pers. comm.). It has also been reported from New Mexico, New York, and Wyoming by W. L. Peterson (1979). This arctic-alpine species is recognized by its erect-spreading leaves, slightly curled when dry, keeled and often weakly undulate above, lanceolate, narrowly acute to acuminate, by its percurrent to shortly excurrent costae, smooth to slightly papillose above, and by its leaf cross section that shows only the abaxial row of cells differentiated, a few 2-stratose marginal cells and the cell walls between lamina cells smooth to slightly bulging. Dicranum acutifolium has often been confused with D. fuscescens but the latter species has a longer, rougher subula that in leaf cross section in the distal half reveals very large and obvious papillae, compared to the smaller and less distinct ones in the former, and margins that are almost completely 2-stratose, while those of D. acutifolium are mostly 1-stratose. It has also been confused with 13. D. brevifolium and for distinctions see discussion thereunder.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

R. R. Ireland (1971b) recorded Dicranum undulatum from Colorado, but W. Weber (pers. comm.) believes this is a misidentification for D. polysetum. Dicranum undulatum is primarily a boreal species that is distinctive because of the yellowish green or yellowish brown, shiny, erect-appressed, lanceolate to oblong-lanceolate, undulate, keeled leaves, the leaf margins that are serrulate to serrate, and broadly recurved in the distal half, the leaf apices that are usually broadly acute, rarely narrowly acute, and the costae that are mostly subpercurrent or sometimes percurrent, smooth to serrulate above on abaxial surface. When the leaf apices are narrow, ± acute, and the costa percurrent, as some D. undulatum plants are on rare occasion, the species when sterile can be confused with D. ontariense. Dicranum undulatum has shiny leaves with twisted apices when dry compared to the more dull leaves that are variously contorted in D. ontariense. When sporophytes are present the aggregate setae (2–5 per perichaetium) of D. ontariense will immediately distinguish it from D. undulatum which has solitary setae. This species recently has been introduced on the University of California Berkeley Campus in a lawn-landscaping area; this is the only California locality where it has been found. The plants are sterile but appear morphologically similar to those growing in native habitats in other parts of North America. Dicranum undulatum of G. F. Weber & C. Mohr (1803) is not a synonym of the taxon treated here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 411. FNA vol. 27, p. 406.
Parent taxa Dicranaceae > Dicranum Dicranaceae > Dicranum
Sibling taxa
D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. undulatum, D. viride
D. acutifolium, D. bonjeanii, D. brevifolium, D. condensatum, D. drummondii, D. elongatum, D. flagellare, D. fragilifolium, D. fulvum, D. fuscescens, D. groenlandicum, D. howellii, D. leioneuron, D. majus, D. montanum, D. muehlenbeckii, D. ontariense, D. pallidisetum, D. polysetum, D. rhabdocarpum, D. scoparium, D. spadiceum, D. spurium, D. tauricum, D. viride
Synonyms D. bergeri var. acutifolium D. bergeri
Name authority (Lindberg & Arnell) C. E. O. Jensen: in H. Weimarck, Förtekn. Skand. Växt., Moss. ed. 2, 18. (1937) Bridel: J. Bot. (Schrader) 1800(2): 294. (1801)
Web links