Dicranum acutifolium |
Dicranum leioneuron |
|
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acuteleaf dicranum moss, sharp-leaf heron's-bill moss |
dicranum moss |
|
Habit | Plants in dense to loose tufts, light green to light brown, dull. | Plants in loose tufts, yellowish green, glossy. |
Stems | 1.5–7 cm, generally branched above, somewhat tomentose with reddish brown rhizoids. |
5–8 cm, scarcely tomentose with whitish to reddish brown rhizoids. |
Leaves | erect-spreading, slightly curled to ± straight when dry, usually with a few undulations in distal part, (3.5–)5–8(–10) mm × 0.5–1 mm, concave below, keeled above, sometimes margins ± involute, lanceolate, narrowly acute to acuminate; margins serrulate to serrate above; laminae 1-stratose or with few 2-stratose regions on or near the distal margins; costa percurrent to shortly excurrent, 1/6–1/4 the width of the leaves at base, smooth to slightly papillose above on abaxial surface, abaxial ridges absent, with a row of guide cells, two stereid bands extending to the apex, adaxial epidermal layer of cells not differentiated, the abaxial layer differentiated; cell walls between lamina cells not or slightly bulging; leaf cells smooth to weakly papillose above on abaxial surface; alar cells 2-stratose, differentiated, often extending to costa; proximal laminal cells linear-rectangular, pitted, (22–)41–60(–90) × (5–)6–8(–12) µm; distal laminal cells short-rectangular, irregularly angled or rounded, not pitted, (7–)12–20(–32) × (4–)8–9(–14) µm. Sexual condition pseudomonoicous; dwarf males on stem rhizoids of female plants; interior perichaetial leaves abruptly short-acuminate, convolute-sheathing. |
erect or spreading, distal leaves with twisted apices when dry, slightly rugose, (3.5–)5–7(–8) × 1–1.5 mm, concave proximally, tubulose distally, from an ovate-lanceolate base to a long or short subula, broadly acute, some leaves short, ovate, somewhat obtuse, forming julaceous regions, in the middle or basal part of the stems; margins not or slightly serrate in the distal part, somewhat involute in the middle part; laminae 1-stratose; costa ending before the apex, 1/13–1/8 the width of the leaves at base, smooth or sometimes the distal part with two weakly serrated ridges on abaxial surface, row of guide cells, two stereid bands, adaxial and abaxial epidermal layers of cells not differentiated or sometimes a few cells in the abaxial layer differentiated; cell walls between lamina cells not bulging; leaf cells smooth; alar cells 2-stratose, differentiated, not extending to costa; proximal laminal cells elongate-sinuose, pitted, (42–)75–87(–107) × (7–)8–10(–16) µm; distal laminal cells short, sinuose, pitted, (16–)30–65(–79) × (7–)9–13(–16) µm. Sexual condition pseudomonoicous; dwarf males on rhizoids of female plants; interior perichaetial leaves abruptly long-acuminate, convolute-sheathing. |
Seta | 1.2–2.5 cm, solitary, yellow to reddish yellow. |
2–4 cm, solitary, yellow, yellowish brown or reddish yellow. |
Capsule | 2–2.7 mm, arcuate, inclined to horizontal, furrowed when dry, rarely slightly strumose, brown to reddish brown; operculum 1.5–2.5 mm. |
2–3.5 mm, arcuate, inclined, smooth, ± furrowed when dry, yellow to yellowish brown; operculum 1.5–2 mm. |
Spores | 14–28 µm. |
14–24 µm. |
Dicranum acutifolium |
Dicranum leioneuron |
|
Phenology | Capsules mature summer. | Capsules mature spring. |
Habitat | Somewhat calcareous soil, boulders, rock outcrops and cliff ledges, sometimes humus, rarely rotten logs | Primarily in hummocks in ombrotrophic and oligotrophic peatlands |
Elevation | 700-2500 m (2300-8200 ft) | 10-1100 m (0-3600 ft) |
Distribution |
AK; ME; MT; NH; NM; NY; WY; AB; BC; MB; NF; NT; NU; ON; QC; YT; Greenland; Europe; Asia |
AK; BC; NB; NF; NS; NT; ON; PE; QC; YT; nw Europe |
Discussion | Dicranum acutifolium has been reported from Colorado by W. A. Weber (1973), but has been deleted from the flora (Weber, pers. comm.). It has also been reported from New Mexico, New York, and Wyoming by W. L. Peterson (1979). This arctic-alpine species is recognized by its erect-spreading leaves, slightly curled when dry, keeled and often weakly undulate above, lanceolate, narrowly acute to acuminate, by its percurrent to shortly excurrent costae, smooth to slightly papillose above, and by its leaf cross section that shows only the abaxial row of cells differentiated, a few 2-stratose marginal cells and the cell walls between lamina cells smooth to slightly bulging. Dicranum acutifolium has often been confused with D. fuscescens but the latter species has a longer, rougher subula that in leaf cross section in the distal half reveals very large and obvious papillae, compared to the smaller and less distinct ones in the former, and margins that are almost completely 2-stratose, while those of D. acutifolium are mostly 1-stratose. It has also been confused with 13. D. brevifolium and for distinctions see discussion thereunder. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Dicranum leioneuron is apparently closely related to D. bonjeanii. It is best distinguished by the subulate leaves with twisted apices when dry, the usual occurrence of julaceous stem portions with broad, short, concave, and somewhat obtuse leaves in contrast to the longer, narrower, subulate, and acute leaves of the other stem portions. Its occurrence only in hummocks in peatlands also is a clue to its identity. The plants rarely produce sporophytes but in any event they cannot be distinguished from those of the other species in section Dicranum. The species probably occurs in peatlands in northeastern United States and it should be looked for in that region. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 411. | FNA vol. 27, p. 404. |
Parent taxa | Dicranaceae > Dicranum | Dicranaceae > Dicranum |
Sibling taxa | ||
Synonyms | D. bergeri var. acutifolium | |
Name authority | (Lindberg & Arnell) C. E. O. Jensen: in H. Weimarck, Förtekn. Skand. Växt., Moss. ed. 2, 18. (1937) | Kindberg: Bull. Torrey Bot. Club 16: 92. (1889) |
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