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dicranoweisia moss, mountain pincushion, mountain thatch-moss

dicranoweisia moss

Habit Plants densely tufted, light to dark green, dull.
Stems

0.5–3 cm, erect, forked, with central strand, with rhizoids confined to base of stems and firmly affixed to substratum.

Leaves

crisped and curled when dry, plane in many leaves, 2-stratose in distal 1/2, cells with longitudinal striolae in distal 1/2 of leaf; usually a few alar cells enlarged on margins, often colored.

lanceolate, flexuose, crisped or curled when dry, 1–3.5 mm;

costa single, subpercurrent, narrow;

margins plane to incurved or recurved, 1- or 2-stratose;

proximal cells elongate, 15–30 µm, smooth;

alar cells with a few enlarged cells on margins, often colored or undifferentiated;

distal cells subquadrate 7–10 µm wide, smooth or with longitudinal cuticular ridges.

Seta

solitary, yellowish, 3–15 mm, smooth.

Sexual condition

autoicous;

pergonial leaves abruptly narrowed to a long or short subula;

perichaetial leaves variable, resembling vegetative leaves or broadly obtuse to acute or short-acuminate.

Capsule

without differentiated annulus;

peristome vertically striolate basally, weakly papillose distally.

light to dark brown when mature, elliptic to short-cylindric, 0.5–2 mm, smooth to longitudinally wrinkled when dry;

annulus present, of 1–3 rows of cells, deciduous or absent;

operculum rostrate, beak straight or oblique;

peristome single, 16 teeth inserted below darkened cells of mouth, teeth entire to divided at apices, red-brown with hyaline tips.

Calyptra

cucullate, smooth, covering 1/3–1/2 of capsule, entire at base.

Spores

spherical, 15–20 µm, weakly papillose.

Specialized

asexual reproduction absent.

asexual reproduction sometimes present, multicellular, on abaxial surface, elliptic to short-linear, of 4–10 cells, 1-seriate or with occasional 2-seriate portions, smooth.

Dicranoweisia crispula

Dicranoweisia

Phenology Capsules mature spring–early summer.
Habitat Forming cushions on siliceous rock or gravel, occasionally epiphytic or epixylic
Elevation 10-2000 m (0-6600 ft)
Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MT; NC; NH; NM; NV; OR; SD; TN; UT; WA; WY; AB; BC; MB; NB; NF; NS; NU; ON; QC; YT; Greenland; Asia
[WildflowerSearch map]
North America; South America; Europe; Asia; Africa; Pacific Islands (Hawaii, New Zealand); Australia; Antarctica
Discussion

Dicranoweisia crispula is an extremely variable species. In stature the plants vary 1–6 cm, while the leaf length varies 1–3.5 mm. The capsules vary considerably from cylindric to short-elliptic, with the length ranging 0.5–2 mm. The shape and sheathing of the perichaetial leaves is also variable. Although this species is autoicous, sex organs are often absent, especially in terrestrial plants, with sporophytes correspondingly infrequent. Differentiation of alar cells is often weak in North American material and the striolation can also be weak, and is best observed on cells adjacent to the costa. This species is widespread mainly at higher elevations in western North America, but sporadic in the eastern part of the continent, again occurring mainly at higher elevations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 20 (2 in the flora).

Species of Dicranoweisia are usually epiphytic, epixylic, epilithic, or terrestrial and are found widespread throughout the world.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf margins widely recurved in many leaves; 1-stratose excepting distal margins, cells smooth; alar cells not differentiated.
D. cirrata
1. Leaf margins plane, erect or incurved to weakly and narrowly recurved; 1-stratose proximally and 2-stratose distally, distal cells usually longitudinally striolate (appearing papillose in transverse section); alar cells sometimes noticeably larger than adjacent cells and often colored.
D. crispula
Source FNA vol. 27, p. 396. FNA vol. 27, p. 395. Author: Wilfred B. Schofield.
Parent taxa Dicranaceae > Dicranoweisia Dicranaceae
Sibling taxa
D. cirrata
Subordinate taxa
D. cirrata, D. crispula
Synonyms Weissia crispula, D. crispula var. compacta, D. contermina, D. roellii, Trichostomum alpinum, Weissia compacta
Name authority (Hedwig) Milde: Bryol. Siles., 49. (1869) Lindberg ex Milde: Bryol. Siles., 48. 1869 ,
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