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fall panicum, fall rosette grass, Wilcox's panicgrass

cypress panicgrass, fork panic grass, fork rosette-panicgrass

Habit Plants cespitose. Plants in small or large clumps, with knotty crowns.
Culms

15-35 cm, stiffly erect, all but the upper 2-4 internodes very short;

nodes glabrous or with weak, reflexed hairs;

internodes purplish-gray, sparsely pubescent;

fall phase developing early, forming erect branches from the lower or midculm nodes, each branch terminating in a partially included panicle of 8-16 spikelets, no sterile shoots formed.

20-100 cm, decumbent to erect, sometimes geniculate;

nodes usually glabrous, sometimes sparsely pilose or densely bearded with retrorse hairs;

internodes often purplish or olive green, lowest internodes usually glabrous, varying to sparsely pubescent;

fall phase usually branching freely, especially from the nodes above the middle, ultimately forming dense, reclining fascicles of divergent branchlets with numerous reduced, thin, often involute blades, secondary panicles often reduced, with few spikelets.

Cauline leaves

usually 3;

sheaths hirsute, hairs papillose-based;

ligules 0.5-1 mm;

blades 4-8 cm long, 2-5 mm wide, all alike, stiffly erect, green to grayish-green, flat, not plicate, sparsely pilose.

4-7;

sheaths usually shorter than the internodes, usually glabrous, occasionally the lower sheaths sparsely to densely soft-pubescent, sheaths of the uppermost leaves sometimes with whitish glandular spots between the prominent veins, margins of all sheaths glabrous or ciliate;

ligules absent or shorter than 1 mm, of hairs;

blades 3.5-14 cm long, 5-14 mm wide, usually thin, distant, spreading to reflexed or (occasionally) ascending, yellow-green to purplish, usually glabrous on both surfaces or (at least the lower blades) more or less densely and softly pubescent, bases constricted (in narrow-bladed subspecies) or narrowly subcordate (in wide-bladed subspecies), margins glabrous or ciliate basally, glabrous distally, blades of the flag leaves usually spreading.

Spikelets

2.4-3.2 mm long, 0.7-1.2 mm wide, ellipsoid to obovoid, often reddish throughout, short-pubescent.

1.5-2.7 mm, usually ellipsoid or obovoid, green or purplish (at least at the base), glabrous or (less commonly) sparsely pubescent or puberulent, often prominently veined, obtuse to acute to beaked.

Lower glumes

0.7-1.2 mm, triangular;

upper glumes and lower lemmas about equaling the upper florets;

upper florets 1.9-2.5 mm, ellipsoid, pointed.

usually less than 1/3 as long as the spikelets, obtuse to acute;

upper glumes usually slightly shorter than or as long as the lower lemmas and upper florets (occasionally extending beyond the floret);

lower florets sterile;

upper florets 1.3-2 mm long, usually less than 1 mm wide, ellipsoid, subacute to obtuse.

Basal

rosettes poorly differentiated;

sheaths glabrous;

blades 2-4 cm, narrow, similar to those of the lower cauline leaves, ascending to spreading.

rosettes well-differentiated;

blades ovate to lanceolate.

Primary

panicles 3-5 cm long, 2-4 cm wide, ovoid, open, shortly exserted, with 12-32 spikelets;

branches short, stiff, spreading;

pedicels mostly 4-8 mm, spreading.

panicles 3-12 cm, long-exserted, usually with many spikelets;

branches wiry, mostly spreading or ascending, usually glabrous, sometimes scabridulous.

2n

= 18.

Dichanthelium wilcoxianum

Dichanthelium dichotomum

Distribution
from FNA
CO; IA; IL; IN; KS; MN; MT; ND; NE; NM; OK; SD; WI; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; PR; NB; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Dichanthelium wilcoxianum grows in dry prairies, especially in sandy or gravelly openings. It is restricted to the Flora region. The primary panicles, which are produced from mid-May to early June, are partially open-pollinated; the secondary panicles, which are produced in June, and occasionally also in September, are cleistogamous.

Some specimens of Dichanthelium oligosanthes subsp. scribnerianum from the southern Great Plains that have prematurely elongating upper internodes resemble D. wilcoxianum, but they have greenish spikelets that are 1.7-2.4 mm wide, an orange spot at the base of the glumes, and larger basal rosettes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dichanthelium dichotomum grows in dry, sandy, clayey, or rocky ground, often in woods, or (more commonly) in moist or wet places, including marshes, bogs, low woods, swamps, and the moist borders of lakes and ponds. Its range extends south from the Flora region into the Caribbean. It is a polymorphic and ubiquitous species, with many of its intergrading subspecies exhibiting traits of other widespread and variable species such as D. commutatum, D. laxiflorum, and D. sphaerocarpon, which often grow at the same sites.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lower nodes hairy.
→ 2
2. Spikelets 1.5-1.8 mm long, upper floret 0.6-0.8 mm wide
subsp. microcarpon
2. Spikelets 1.8-2.5 mm long; upper floret 0.7-1.0 mm wide.
→ 3
3. Spikelets usually glabrous; midculm blades usually 5-7 mm wide
subsp. dichotomum
3. Spikelets pubescent; midculm blades usually 7-14 mm wide.
→ 4
4. Lower sheaths and blades glabrous or sparsely pubescent
subsp. nitidum
4. Lower sheaths and blades more or less densely velvety pubescent
subsp. mattamuskeetense
1. Lower nodes glabrous.
→ 5
5. Larger blades more than 1 cm wide; sheaths often with pale glandular spots between the prominent veins; spikelets 1.9-2.6 mm long, acute to beaked
subsp. yadkinense
5. Larger blades less than 1 cm wide; sheaths without glandular spots; spikelets 1.5-2.3 mm long, obtuse to subacute.
→ 6
6. Culms weak, ultimately reclining or sprawling, often flattened
subsp. lucidum
6. Culms erect, terete.
→ 7
7. Blades usually spreading; spikelets ellipsoid, 1.8-2.3 mm long, rarely purplish at the base
subsp. dichotomum
7. Blades usually ascending or erect; spikelets broadly ellipsoid or obovoid, 1.5-1.8 mm long, often purplish at the base
subsp. roanokense
Source FNA vol. 25, p. 449. FNA vol. 25, p. 432.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Dichanthelium > sect. Linearifolia Poaceae > subfam. Panicoideae > tribe Paniceae > Dichanthelium > sect. Dichanthelium
Sibling taxa
D. aciculare, D. acuminatum, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. dichotomum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. portoricense, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. strigosum, D. tenue, D. wrightianum, D. ×anthophysum
D. aciculare, D. acuminatum, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. portoricense, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. strigosum, D. tenue, D. wilcoxianum, D. wrightianum, D. ×anthophysum
Subordinate taxa
D. dichotomum subsp. dichotomum, D. dichotomum subsp. lucidum, D. dichotomum subsp. mattamuskeetense, D. dichotomum subsp. microcarpon, D. dichotomum subsp. nitidum, D. dichotomum subsp. roanokense, D. dichotomum subsp. yadkinense
Synonyms Panicum wilcoxianum Panicum niditum, Panicum dichotomum var. barbulatum, Panicum dichotomum
Name authority (Vasey) Freckmann (L.) Gould
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