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cushion-tuft panicgrass, roughhair rosette grass

cypress panicgrass, fork panic grass, fork rosette-panicgrass

Habit Plants densely cespitose. Plants in small or large clumps, with knotty crowns.
Culms

5-45 cm, slender, erect or spreading;

from a dense tuft of predominantly basal leaves, lower internodes short, upper 3-5 internodes elongate;

nodes glabrous or bearded;

internodes glabrous or pilose;

fall phase with spreading culms and branches arising from near the bases forming a dense, flat tuft.

20-100 cm, decumbent to erect, sometimes geniculate;

nodes usually glabrous, sometimes sparsely pilose or densely bearded with retrorse hairs;

internodes often purplish or olive green, lowest internodes usually glabrous, varying to sparsely pubescent;

fall phase usually branching freely, especially from the nodes above the middle, ultimately forming dense, reclining fascicles of divergent branchlets with numerous reduced, thin, often involute blades, secondary panicles often reduced, with few spikelets.

Cauline leaves

2-4;

lower cauline sheaths longer than the internodes, mostly glabrous or pilose with ascending hairs, margins finely ciliate;

ligules 0.2-2 mm, at low magnification appearing to be membranous and ciliate, at high magnification evidently of hairs that are coherent at the base;

blades 1.5-6 cm long, 3-8 mm wide, lanceolate, glabrous or softly pilose, margins with prominent papillose-based cilia, at least basally.

4-7;

sheaths usually shorter than the internodes, usually glabrous, occasionally the lower sheaths sparsely to densely soft-pubescent, sheaths of the uppermost leaves sometimes with whitish glandular spots between the prominent veins, margins of all sheaths glabrous or ciliate;

ligules absent or shorter than 1 mm, of hairs;

blades 3.5-14 cm long, 5-14 mm wide, usually thin, distant, spreading to reflexed or (occasionally) ascending, yellow-green to purplish, usually glabrous on both surfaces or (at least the lower blades) more or less densely and softly pubescent, bases constricted (in narrow-bladed subspecies) or narrowly subcordate (in wide-bladed subspecies), margins glabrous or ciliate basally, glabrous distally, blades of the flag leaves usually spreading.

Spikelets

1.1-2.1 mm, obovoid to broadly ellipsoid, glabrous or pubescent, hairs not papillose-based.

1.5-2.7 mm, usually ellipsoid or obovoid, green or purplish (at least at the base), glabrous or (less commonly) sparsely pubescent or puberulent, often prominently veined, obtuse to acute to beaked.

Lower glumes

1/3 - 1/2 as long as the spikelets, acute to obtuse;

upper florets 0.8-1.7 mm, ellipsoid, subacute.

usually less than 1/3 as long as the spikelets, obtuse to acute;

upper glumes usually slightly shorter than or as long as the lower lemmas and upper florets (occasionally extending beyond the floret);

lower florets sterile;

upper florets 1.3-2 mm long, usually less than 1 mm wide, ellipsoid, subacute to obtuse.

Basal

rosettes poorly differentiated;

blades 1-5 cm, lanceolate, grading into the cauline blades.

rosettes well-differentiated;

blades ovate to lanceolate.

Primary

panicles short- to long-exserted;

rachises and branches often pilose.

panicles 3-12 cm, long-exserted, usually with many spikelets;

branches wiry, mostly spreading or ascending, usually glabrous, sometimes scabridulous.

Dichanthelium strigosum

Dichanthelium dichotomum

Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TN; TX; VA; PR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; PR; NB; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Dichanthelium strigosum extends from the southeastern Flora region south into Mexico, the Caribbean, and into northern South America.

The primary panicles are briefly open-pollinated in April or May; the secondary panicles, which are produced from May through November, are cleistogamous. The three subspecies are mostly sympatric and sometimes grow together, with occasional intergradation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dichanthelium dichotomum grows in dry, sandy, clayey, or rocky ground, often in woods, or (more commonly) in moist or wet places, including marshes, bogs, low woods, swamps, and the moist borders of lakes and ponds. Its range extends south from the Flora region into the Caribbean. It is a polymorphic and ubiquitous species, with many of its intergrading subspecies exhibiting traits of other widespread and variable species such as D. commutatum, D. laxiflorum, and D. sphaerocarpon, which often grow at the same sites.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets pubescent, broadly ellipsoid, 1.6-2.1 mm long; lower glumes about 1/2 as long as the spikelets; blades glabrous
subsp. leucoblepharis
1. Spikelets glabrous, obovoid, 1.1-1.8 mm long; lower glumes about 1/3 as long as the spikelets; blades pilose or glabrous.
→ 2
2. Blades pilose; spikelets 1.1-1.6 mm long
subsp. strigosum
2. Blades glabrous or sparsely pilose near the base; spikelets 1.4-1.8 mm long
subsp. glabrescens
1. Lower nodes hairy.
→ 2
2. Spikelets 1.5-1.8 mm long, upper floret 0.6-0.8 mm wide
subsp. microcarpon
2. Spikelets 1.8-2.5 mm long; upper floret 0.7-1.0 mm wide.
→ 3
3. Spikelets usually glabrous; midculm blades usually 5-7 mm wide
subsp. dichotomum
3. Spikelets pubescent; midculm blades usually 7-14 mm wide.
→ 4
4. Lower sheaths and blades glabrous or sparsely pubescent
subsp. nitidum
4. Lower sheaths and blades more or less densely velvety pubescent
subsp. mattamuskeetense
1. Lower nodes glabrous.
→ 5
5. Larger blades more than 1 cm wide; sheaths often with pale glandular spots between the prominent veins; spikelets 1.9-2.6 mm long, acute to beaked
subsp. yadkinense
5. Larger blades less than 1 cm wide; sheaths without glandular spots; spikelets 1.5-2.3 mm long, obtuse to subacute.
→ 6
6. Culms weak, ultimately reclining or sprawling, often flattened
subsp. lucidum
6. Culms erect, terete.
→ 7
7. Blades usually spreading; spikelets ellipsoid, 1.8-2.3 mm long, rarely purplish at the base
subsp. dichotomum
7. Blades usually ascending or erect; spikelets broadly ellipsoid or obovoid, 1.5-1.8 mm long, often purplish at the base
subsp. roanokense
Source FNA vol. 25, p. 446. FNA vol. 25, p. 432.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Dichanthelium > sect. Strigosa Poaceae > subfam. Panicoideae > tribe Paniceae > Dichanthelium > sect. Dichanthelium
Sibling taxa
D. aciculare, D. acuminatum, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. dichotomum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. portoricense, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. tenue, D. wilcoxianum, D. wrightianum, D. ×anthophysum
D. aciculare, D. acuminatum, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. portoricense, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. strigosum, D. tenue, D. wilcoxianum, D. wrightianum, D. ×anthophysum
Subordinate taxa
D. strigosum subsp. glabrescens, D. strigosum subsp. leucoblepharis, D. strigosum subsp. strigosum
D. dichotomum subsp. dichotomum, D. dichotomum subsp. lucidum, D. dichotomum subsp. mattamuskeetense, D. dichotomum subsp. microcarpon, D. dichotomum subsp. nitidum, D. dichotomum subsp. roanokense, D. dichotomum subsp. yadkinense
Synonyms Panicum strigosum Panicum niditum, Panicum dichotomum var. barbulatum, Panicum dichotomum
Name authority (Muhl. ex Elliott) Freckmann (L.) Gould
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