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blunt-glumed panicgrass

hairy panic grass, hairy perennial panicgrass, hairy rosette-panicgrass, hotsprings rosette grass, panic laineux, tapered rosette grass, western witchgrass, woolly panic grass, woolly witchgrass

Habit Plants usually densely cespitose. Plants more or less densely cespitose.
Culms

15-50 cm, slender, wiry;

internodes olive green to purplish, densely puberulent or glabrous;

fall phase spreading or decumbent, branching extensively from the lower and midculm nodes, producing numerous congested fascicles of reduced, flat or involute blades and reduced secondary panicles.

15-100 cm (rarely taller), usually thicker than 1 mm, weak and wiry or relatively stout and rigid, erect, ascending or decumbent;

nodes occasionally swollen, glabrous or densely pubescent, often with a glabrous or viscid ring below;

internodes purplish or olive green or grayish-green, to yellowish-green, variously pubescent, with hairs of 2 lengths or glabrous;

fall phase erect, spreading, or decumbent, usually branching extensively at all but the uppermost nodes, ultimately forming dense fascicles of branchlets with reduced, flat or involute blades and reduced secondary panicles with few spikelets.

Cauline leaves

4-7;

sheaths much shorter than the internodes, densely crisp-puberulent, velvety-puberulent, or glabrous, often ciliate along the margins;

ligules shorter than 0.5 mm;

blades 2-7 cm long (seldom longer), 2.5-8 mm wide (rarely wider), spreading, firm, flat or slightly involute, without prominently raised veins, not longitudinally wrinkled, densely puberulent or glabrous abaxially, glabrous, sparsely puberulent, or pubescent adaxially, bases subcordate, with papillose-based cilia, margins often whitish and scabridulous.

4-7;

sheaths usually shorter than the internodes, glabrous or densely and variously pubescent with hairs shorter than 3 mm, margins ciliate or glabrous;

ligules and pseudoligules 1-5 mm, of hairs;

blades 2-12 cm long (rarely longer), 2-12 mm wide (rarely wider), firm or lax, spreading to reflexed or stiffly ascending, yellowish-green or grayish-green to olivaceous, densely to sparsely and variously pubescent, margins similar or occasionally whitish-scabridulous, margins often with papillose-based cilia, at least basally, bases rounded or subcordate.

Spikelets

1.5-2.6 mm, obovoid-pyriform, planoconvex in side view, puberulent, pubescent, or glabrous, attenuate basally, apices usually broadly rounded.

1.1-2.1 mm, obovoid to ellipsoid, yellowish-green to olivaceous or purplish, variously pubescent, obtuse or subacute.

Lower glumes

0.6-1.4 mm, thin, weakly-veined, attached about 0.2 mm below the upper glumes, clasping at the base;

upper glumes as long as or slightly shorter than the lower lemmas;

upper florets 1.4-2 mm, broadly ellipsoid, apices subacute, minutely puberulent.

usually 1/4 - 1/2 as long as the spikelets, obtuse to acute;

upper glumes and lower lemmas subequal, equaling the upper florets at maturity, or occasionally the upper glumes slightly shorter, not strongly veined;

lower florets sterile;

upper florets 1.1-1.7 mm long, 0.6-1 mm wide, ellipsoid, obtuse to acute or minutely umbonate or apiculate.

Basal

rosettes well-differentiated;

blades 1.5-6 cm, ovate to lanceolate.

rosettes usually well-differentiated;

blades ovate to lanceolate.

Primary

panicles 2-7 cm long, 2/3 to nearly as wide as long, with relatively few spikelets, exserted;

branches flexuous, spreading or reflexed, scabridulous to densely puberulent.

panicles 3-12 cm, 1/4 - 3/4 as wide as long, usually open, well-exserted, rather dense;

rachises glabrous, puberulent, or more or less densely pilose, at least basally.

2n

= 18.

= 18.

Dichanthelium portoricense

Dichanthelium acuminatum

Distribution
from USDA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; AB; BC; MB; NB; NS; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Dichanthelium portoricense grows in sandy woods, low pinelands, savannahs, and coastal sand dunes, usually in moist places. Its range extends south from the Flora region into Mexico, the Caribbean, and Mesoamerica. It is a highly variable species with numerous intergrading forms, some possibly resulting from hybridization with other widespread species in the same region, such as D. sphaerocarpon and D. commutatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dichanthelium acuminatum is common and ubiquitous in dry to wet, open, sandy or clayey woods, clearings, bogs, and swamps, or in saline soil near hot springs, growing in much of the Flora region and extending into northern South America. It is probably the most polymorphic and troublesome species in the genus. The treatment presented here attempts to delimit the major variants present, but does not fully reflect the intricate reticulate pattern of morphological variation that exists. There is considerable overlap among the nine subspecies recognized and, in addition, there appears to be widespread introgression from other Dichanthelium species, such as D. dichotomum, D. sphaerocarpon, D. ovale, and D. aciculare into the D. acuminatum complex, contributing to the taxonomic difficulties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets 1.8-2.6 mm long, usually densely pubescent or puberulent (rarely glabrous); cauline blades 4-7 cm long, 3.5-8 mm wide
subsp. patulum
1. Spikelets 1.5-2.0 mm long, puberulent to nearly glabrous; cauline blades 2-5 cm long, 2.5-4.5 mm wide
subsp. portoricense
1. Lower portion of the culms and lower sheaths usually glabrous or sparsely pubescent.
→ 2
2. Primary panicles congested, more than twice as long as wide; spikelets ascending to appressed
subsp. spretum
2. Primary panicles open, less than twice as long as wide; spikelets diverging to ascending.
→ 3
3. Blades green or purplish, the margins not conspicuously ciliate at the base; spikelets 1.1-1.5 mm long, usually ellipsoid
subsp. longiligulatum
3. Blades often yellowish-green, the margins usually with long, papillose-based cilia at the base; spikelets 1.3-1.6 mm long, usually obovoid
subsp. lindheimeri
1. Lower portion of the culms and lower sheaths densely and variously pubescent or puberulent.
→ 4
4. Culms 15-30 cm tall; midculm sheaths nearly as long as the internodes; blades usually 2-6.5 cm long, less than 8 times longer than wide
subsp. sericeum
4. Culms usually 30-100 cm tall; midculm sheaths about 1/2 as long as the internodes; blades usually 6-12 cm long, more than 8 times longer than wide.
→ 5
5. Culms and lower sheaths densely covered with spreading, villous hairs or soft, thin, papillose-based hairs, often with shorter hairs underneath; blades softly pubescent to velvety on the abaxial surfaces.
→ 6
6. Primary panicles usually poorly exserted, on peduncles less than 6 cm long; blades suberect, the margins lacking cilia on the distal 1/2
subsp. thermale
6. Primary panicles usually well-exserted, on peduncles more than 8 cm long; blades ascending to spreading, the margins ciliate along most of their length
subsp. acuminatum
5. Culms and sheaths pilose with papillose-based hairs to hispid, with mostly ascending hairs, or densely puberulent with a few longer, ascending hairs also present; blades appressed-pubescent or puberulent abaxially, not velvety to the touch.
→ 7
7. Sheaths and culms densely puberulent, scattered long hairs often present also
subsp. columbianum
7. Sheaths and culms pilose with papillose-based hairs, the hairs mostly ascending, occasionally with inconspicuous, shorter hairs underneath.
→ 8
8. Blades usually 6-12 mm wide, spreading to ascending, the adaxial surfaces nearly glabrous or with hairs shorter than 3 mm long; spikelets 1.5-2 mm
subsp. fasciculatum
8. Blades usually 2-6 mm wide, erect to ascending, spreading or reflexed, the adaxial surfaces glabrous or with hairs 3-6 mm long; spikelets 1.1-1.6 mm long.
→ 9
9. Blades erect to ascending, the adaxial surfaces long-pilose; spikelets 1.3-1.6 mm long, usually broadly obovoid
subsp. implicatum
9. Blades ascending, spreading, or reflexed, the adaxial surfaces glabrous or sparsely pubescent; spikelets 1.1-1.5 mm long, usually ellipsoid
subsp. leucothrix
Source FNA vol. 25, p. 441. FNA vol. 25, p. 422.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Dichanthelium > sect. Lancearia Poaceae > subfam. Panicoideae > tribe Paniceae > Dichanthelium > sect. Lanuginosa
Sibling taxa
D. aciculare, D. acuminatum, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. dichotomum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. strigosum, D. tenue, D. wilcoxianum, D. wrightianum, D. ×anthophysum
D. aciculare, D. boreale, D. boscii, D. chamaelonche, D. clandestinum, D. commutatum, D. consanguineum, D. depauperatum, D. dichotomum, D. ensifolium, D. erectifolium, D. latifolium, D. laxiflorum, D. leibergii, D. linearifolium, D. malacophyllum, D. nodatum, D. nudicaule, D. oligosanthes, D. ovale, D. pedicellatum, D. perlongum, D. polyanthes, D. portoricense, D. ravenelii, D. scabriusculum, D. scoparium, D. sphaerocarpon, D. strigosum, D. tenue, D. wilcoxianum, D. wrightianum, D. ×anthophysum
Subordinate taxa
D. portoricense subsp. patulum, D. portoricense subsp. portoricense
D. acuminatum subsp. acuminatum, D. acuminatum subsp. columbianum, D. acuminatum subsp. fasciculatum, D. acuminatum subsp. implicatum, D. acuminatum subsp. leucothrix, D. acuminatum subsp. lindheimeri, D. acuminatum subsp. longiligulatum, D. acuminatum subsp. sericeum, D. acuminatum subsp. spretum, D. acuminatum subsp. thermale
Synonyms Panicum portoricense Panicum acuminatum, Panicum lanuginosum
Name authority (Desv. ex Ham.) B.E Hansen & Wunderlin (Sw) Gould & C.A. Clark
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