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few flower dicentra, few-flower bleeding-heart, shorthorn steer's-head

Oregon bleeding heart, Pacific bleeding-heart, Pacific bleedinghearts, western bleeding-heart

Habit Plants perennial, scapose, from rhizomes or clusters of spindle-shaped tubers, bulblets often present at proximal ends of tubers or along rhizomes. Plants perennial, scapose, from elongate, stout rhizomes.
Leaves

(7-)9-13(-16) × 3-7(-10) cm;

petiole (2-)4-7(-10) cm;

blade with 3-4 orders of leaflets and lobes; ultimate lobes linear-lanceolate, (2-)7-13(-18) × 1.5-3 mm, occasionally irregular, minutely apiculate.

(15-)25-40(-55) × (8-)12-20(-35) cm;

blade with 3-5 orders of leaflets and lobes;

abaxial surface and sometimes adaxial surface glaucous; penultimate lobes oblong, distal ones usually coarsely 3-toothed at apex, (4-)10-20(-50) × (1.5-)3-4(-8) mm.

Inflorescences

racemose, 1-3-flowered, barely exceeding leaves;

bracts ovate, 4-5 × 2-3 mm.

paniculate, 2-30-flowered, usually exceeding leaves;

bracts linear-lanceolate, 4-7(-12) × 1-2 mm, apex acuminate.

Flowers

erect to nodding;

pedicels 5-25 mm;

sepals ovate to lanceolate, 5-8 × 2-4 mm;

outer petals white to pink, (15-)18-22(-25) × 3-6 mm, reflexed portion (5-)7-8(-11) mm;

inner petals purple, (15-)18-22(-24) mm, blade spoon-shaped, 2-3 mm, claw obovate-elliptic, ca. 10 × 3-4 mm, crest absent;

filaments of each bundle connate at base and near apex, distinct in between, distinct portion of median filament forming loop that almost doubles back to its proximal end; nectariferous tissue borne at lowermost point of loop and often extending to base of median filament;

style 7-11 mm;

stigma 2-lobed, much reduced, ca. 2 times wider than style.

pendent;

sepals lanceolate to ovate or nearly round, 2-7 × 2-3 mm;

petals rose-purple, pink, cream, or pale yellow, rarely white;

outer petals (12-)16-19(-24) × 3-6 mm, reflexed portion 2-5 mm;

inner petals (12-)15-18(-22) mm, blade 2-4 mm wide, claw linear-elliptic to linear-lanceolate, 7-10(-12) × 1-2 mm, crest 1-2 mm diam., exceeding apex by 1-2 mm;

filaments of each bundle connate from base to shortly below anthers except for a 2-3 mm portion of median filament just above base; nectariferous tissue borne along distinct portion of median filament;

style 3-9 mm;

stigma rhomboid, 2-horned.

Capsules

spindle-shaped to ovoid, 10-15 × 4-6 mm.

oblong, 4-5 mm diam.

Seeds

reniform, ca. 2 mm diam., smooth, elaiosome present.

reniform, ca. 2 mm diam., finely reticulate, elaiosome present.

2n

= 16.

Dicentra pauciflora

Dicentra formosa

Phenology Flowering late spring–late summer.
Habitat Openings in coniferous forests, in volcanic and granitic soils
Elevation 1200-2700 m (3900-8900 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Andrews has been cited almost universally as the author of Fumaria formosa. However, Haworth's authorship of the sixth volume of Andrews' Botanists' Repository (in which this species was originally described) generally has been overlooked, and it was actually Haworth who first delineated F. formosa (W. T. Stearn 1944).

Early attempts to cross Dicentra formosa with D. eximia (2n = 16) failed, possibly because the D. formosa parents were tetraploids. Several later hybrids between the two species received plant patents and have become widely marketed throughout the flora area and elsewhere (K. R. Stern 1961, 1968; K. R. Stern and M. Ownbey 1971).

Both subspecies, as well as hybrids between them and Dicentra eximia, are widely cultivated.

The Skagit used a decoction of the roots of Dicentra formosa to expel worms; they chewed raw roots for toothaches (D. E. Moerman 1986, species not indicated).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals rose-purple to pink, rarely white; leaf blades adaxially not glaucous (rarely glaucescent).
subsp. formosa
1. Petals cream-colored or rarely pale yellow, rose-tipped; leaf blades adaxially distinctly glaucous.
subsp. oregana
Source FNA vol. 3. FNA vol. 3.
Parent taxa Fumariaceae > Dicentra Fumariaceae > Dicentra
Sibling taxa
D. canadensis, D. chrysantha, D. cucullaria, D. eximia, D. formosa, D. nevadensis, D. ochroleuca, D. uniflora
D. canadensis, D. chrysantha, D. cucullaria, D. eximia, D. nevadensis, D. ochroleuca, D. pauciflora, D. uniflora
Subordinate taxa
D. formosa subsp. formosa, D. formosa subsp. oregana
Synonyms Fumaria formosa, D. saccata
Name authority S. Watson: Bot. California 2: 429. (1880) (Haworth) Walpers: Repert. Bot. Syst. 1: 118. (1842)
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