Dicentra formosa |
Dicentra formosa subsp. formosa |
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Oregon bleeding heart, Pacific bleeding-heart, Pacific bleedinghearts, western bleeding-heart |
bleeding heart, Pacific bleeding-heart |
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Habit | Plants perennial, scapose, from elongate, stout rhizomes. | |||||
Leaves | (15-)25-40(-55) × (8-)12-20(-35) cm; blade with 3-5 orders of leaflets and lobes; abaxial surface and sometimes adaxial surface glaucous; penultimate lobes oblong, distal ones usually coarsely 3-toothed at apex, (4-)10-20(-50) × (1.5-)3-4(-8) mm. |
blades abaxially glaucous, adaxially not glaucous (rarely glaucescent). |
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Inflorescences | paniculate, 2-30-flowered, usually exceeding leaves; bracts linear-lanceolate, 4-7(-12) × 1-2 mm, apex acuminate. |
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Flowers | pendent; sepals lanceolate to ovate or nearly round, 2-7 × 2-3 mm; petals rose-purple, pink, cream, or pale yellow, rarely white; outer petals (12-)16-19(-24) × 3-6 mm, reflexed portion 2-5 mm; inner petals (12-)15-18(-22) mm, blade 2-4 mm wide, claw linear-elliptic to linear-lanceolate, 7-10(-12) × 1-2 mm, crest 1-2 mm diam., exceeding apex by 1-2 mm; filaments of each bundle connate from base to shortly below anthers except for a 2-3 mm portion of median filament just above base; nectariferous tissue borne along distinct portion of median filament; style 3-9 mm; stigma rhomboid, 2-horned. |
petals rose-purple to pink, rarely white. |
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Capsules | oblong, 4-5 mm diam. |
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Seeds | reniform, ca. 2 mm diam., finely reticulate, elaiosome present. |
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2n | = 16, 32. |
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Dicentra formosa |
Dicentra formosa subsp. formosa |
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Phenology | Flowering early spring–early fall. | |||||
Habitat | Loam or gravel soils in moist woods and clearings, and along banks of streams | |||||
Elevation | 0-2250 m (0-7400 ft) | |||||
Distribution |
CA; OR; WA; BC
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CA; OR; WA; BC |
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Discussion | Subspecies 2 (2 in the flora). Andrews has been cited almost universally as the author of Fumaria formosa. However, Haworth's authorship of the sixth volume of Andrews' Botanists' Repository (in which this species was originally described) generally has been overlooked, and it was actually Haworth who first delineated F. formosa (W. T. Stearn 1944). Early attempts to cross Dicentra formosa with D. eximia (2n = 16) failed, possibly because the D. formosa parents were tetraploids. Several later hybrids between the two species received plant patents and have become widely marketed throughout the flora area and elsewhere (K. R. Stern 1961, 1968; K. R. Stern and M. Ownbey 1971). Both subspecies, as well as hybrids between them and Dicentra eximia, are widely cultivated. The Skagit used a decoction of the roots of Dicentra formosa to expel worms; they chewed raw roots for toothaches (D. E. Moerman 1986, species not indicated). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
This subspecies occurs in two chromosomal races: tetraploids (2n = 32), distributed from the Cascade Mountains of Oregon southward throughout the Coast Ranges to central California, and diploids (n = 16), distributed from Vancouver Island and British Columbia southward through the Cascades and Coast Ranges, and along the western slopes of the Sierra Nevada to Monterey and Tulare counties, California. The flowers of both races vary appreciably in color, shape, and size. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 3. | FNA vol. 3. | ||||
Parent taxa | Fumariaceae > Dicentra | Fumariaceae > Dicentra > Dicentra formosa | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Fumaria formosa, D. saccata | D. formosa var. breviflora | ||||
Name authority | (Haworth) Walpers: Repert. Bot. Syst. 1: 118. (1842) | unknown | ||||
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