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big-head rabbit-tobacco, bighead pygmycudweed

many-stem rabbit-tobacco, spring pygmycudweed, spring rabbit-tobacco

Habit Plants grayish green to silvery, 3–15 cm, sericeous to lanuginose. Plants greenish to grayish, 2–15(–25) cm, ± lanuginose.
Stems

mostly 2–10;

branches proximal and distal (distal opposite or, sometimes, appearing alternate when unequal), rarely none.

mostly 2–10;

branches proximal and distal (distal subopposite), rarely none.

Leaves

largest 7–15 × 2–4 mm;

capitular leaves subtending glomerules, also visible between and surpassing heads.

largest 7–13 × 2–4 mm;

capitular leaves subtending glomerules, also ± hidden between and surpassed by heads.

Receptacles

broadly or narrowly conic, 0.4–0.6 mm or ± 0.9–1.1 mm, heights 0.5–0.7 or 2–2.4 times diams.

pulvinate, 0.3–0.6 mm, heights ± 0.2–0.5 times diams.

Bisexual florets

0.

0.

Heads

in strictly dichasiform or pseudo-polytomous arrays (sometimes appearing monochasiiform), cylindric to ± ellipsoid, 3.5–4.5 mm, heights 2–3 times diams.

mostly distal, in subdichasiform arrays, campanulate to ± spheric, 2–3.3 mm, heights ± equal to diams.

Cypselae

± angular, obcompressed, mostly 0.9–1.2 mm.

rounded, ± terete, mostly 0.7–0.9 mm.

Pistillate

paleae imbricate, longest 2.5–4 mm.

paleae scarcely imbricate, longest 1.9–2.7 mm.

Staminate

paleae ± 3, apices erect to somewhat spreading, ± plane.

paleae mostly 3–5, apices somewhat spreading, ± plane.

Functionally

staminate florets 2–4;

ovaries partly developed, 0.4–0.6 mm;

corollas hidden in heads, actinomorphic, 1.4–2 mm, glabrous, lobes equal.

staminate florets 3–5;

ovaries vestigial, 0–0.1 mm;

corollas hidden in heads, actinomorphic, 1.8–2.5 mm, often ± spreading-arachnoid, lobes equal.

2n

= 26.

Diaperia prolifera

Diaperia verna

Distribution
from FNA
AL; AR; CO; KS; LA; MO; MS; MT; NE; NM; OK; SD; TX; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; GA; LA; NM; OK; SC; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Intermediates between the two varieties of Diaperia prolifera occur where their ranges meet in central Texas and central Oklahoma. The strictly dichasiform or pseudo-polytomous branching pattern of D. prolifera is distinctive and diagnostic within the genus. Specimens of D. prolifera from introductions around a wool mill in South Carolina (G. L. Nesom 2004c, as Evax prolifera) are as yet undetermined to variety and are not included in the distributions below.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The two varieties of Diaperia verna intergrade within a broad band inland from the Gulf of Mexico in southeastern Texas. Though some specimens are difficult to assign with confidence, the varieties show enough correlated geographic and ecologic segregation to warrant taxonomic recognition.

As neotypified by J. D. Morefield (2004), the name Evax verna now applies to the taxon that de Candolle named E. multicaulis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants grayish to greenish, loosely lanuginose; heads 4–40+ in largest glomerules; receptacle heights mostly 0.5–0.7 times diams.; capitular leaves usually ± spreading, scarcely involucral, not or scarcely carinate, pliant to somewhat rigid; distal branches mostly spreading to ascending; longest pistillate paleae 3.3–4 mm
var. prolifera
1. Plants silvery white, tightly sericeous; heads borne singly, or 2–3 in largest glomerules; receptacle heights mostly 2–2.4 times diams.; capitular leaves erect, involucral, proximally carinate, becoming indurate; distal branches strictly ascending to erect; longest pistillate paleae 2.5–3.2 mm
var. barnebyi
1. Pistillate paleae collectively hidden by thick lanuginose indument; heads ± campanulate, larg- est mostly 2–2.5 mm
var. verna
1. Pistillate paleae individually visible through thin sericeous indument; heads ± spheric, largest mostly 2.5–3.3 mm
var. drummondii
Source FNA vol. 19, p. 462. FNA vol. 19, p. 461.
Parent taxa Asteraceae > tribe Gnaphalieae > Diaperia Asteraceae > tribe Gnaphalieae > Diaperia
Sibling taxa
D. candida, D. verna
D. candida, D. prolifera
Subordinate taxa
D. prolifera var. barnebyi, D. prolifera var. prolifera
D. verna var. drummondii, D. verna var. verna
Synonyms Evax prolifera Evax verna, Evax multicaulis
Name authority (Nuttall ex de Candolle) Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 338. (1840) (Rafinesque) Morefield: Novon 14: 468. (2004)
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