Desmodium retinens |
Desmodium triflorum |
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Santa Rita Mountain ticktrefoil |
creeping tick trefoil, matty desmodium, three-flower beggarweed, threeflower ticktrefoil, tropical trefoil |
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Habit | Herbs, perennial, diffuse; base woody, rootstock woody. | Herbs, annual or perennial, often mat-forming; stoloniferous. |
Stems | erect, ascending, or procumbent, usually striate, 30–60(–100) cm, sparsely to densely pilose and uncinate-pubescent. |
prostrate, densely branched, 20–80 cm, ascending-pilose or strigose. |
Leaves | trifoliolate, sometimes unifoliolate proximally; stipules persistent, recurved, subulate, 2–3 mm; petiole 7–25 mm; leaflet blades usually oblong-ovate to narrowly so, sometimes broadly elliptic to oblong, apex obtuse or acute, surfaces sparsely uncinate-puberulent; terminal blade 10–30(–35) × 8–10 mm, length 1–3.5 times width. |
trifoliolate; stipules persistent, narrowly ovate, 3–5 mm; petiole 3–7.5 mm; leaflet blades broadly obovate or cuneate-obovate, often folding downwards, apex emarginate, surfaces usually sparsely uncinate-puberulent or subappressed-pilose along midrib abaxially, rarely entire surface, glabrous adaxially; terminal blade 5–10 × 3–11 mm, length 0.8–1.2 times width. |
Inflorescences | usually unbranched, sometimes branched; rachis patent uncinate-puberulent; primary bracts narrowly ovate, 2.5–3 mm. |
terminal fascicles opposite distal leaf, appearing axillary, unbranched; rachis pubescent; primary bracts narrowly ovate, 4 mm. |
Pedicels | 8–15 mm. |
8–12 mm. |
Flowers | calyx 1.5–2.5 mm, pubescent, tube 1 mm; abaxial lobes 1.5 mm, lateral lobes 1 mm; corolla pink-purple, 4–5 mm. |
calyx 2.5–3 mm, appressed-pubescent, tube 1.2 mm; abaxial lobes 1.5–1.7 mm, lateral lobes 1.5–1.7 mm, adaxial lobes deeply 2-toothed; corolla pale pink to purplish, 4–5 mm, keel distinctly longer than wings. |
Loments | sutures subequally crenate; connections central, 1/5–1/4 as broad as segments; segments 2–7, rounded or broadly elliptic, 2.5–3 × 2.5 mm, abaxially and adaxially rounded, glabrous or slightly puberulent; stipe 1–2 mm. |
sutures weakly crenate abaxially, with broad connection between segments, barely sinuate adaxially; connections 2/3–4/5 as broad as segments; segments 3–5, nearly square, 2.5–3.5 × 2.5–3 mm, symmetrically convex abaxially, slightly concave adaxially, inconspicuously uncinate-pubescent or glabrescent; stipe 0 mm. |
2n | = 22. |
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Desmodium retinens |
Desmodium triflorum |
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Phenology | Flowering late summer–fall. | Flowering year-round. |
Habitat | Wooded or grassy rocky slopes, oak or oak-pine forests. | Ruderal areas, lawns, disturbed open woodlands. |
Elevation | 1400–2100 m. (4600–6900 ft.) | 0–50 m. (0–200 ft.) |
Distribution |
AZ; Mexico; Central America (Guatemala) |
FL; LA; Central America; South America; Mexico (Jalisco, Sinaloa); West Indies; Asia; Africa [Introduced in North America; introduced also in Indian Ocean Islands, Pacific Islands, Australia]
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Discussion | Desmodium retinens is known in the flora area from Cochise, Graham, Pima, and Santa Cruz counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Inflorescences of Desmodium triflorum are produced opposite the leaf at the distal end of stem and are usually described as axillary. The branching system of D. triflorum is a monopodial sympodium and the inflorescences are terminal (H. Ohashi and T. Nemoto 1986). Evolution of inflorescences in Desmodium is inferred from comparative morphology and anatomy with Campylotropis, Kummerowia, and Lespedeza (Nemoto and Ohashi 1990, 1993, 1996). Based on molecular and morphological data, H. Ohashi and K. Ohashi (2018) transferred Desmodium triflorum and the other species formerly in Desmodium sects. Nicolsonia (de Candolle) Bentham and Sagotia (Duchassaing & Walpers) Bentham to the genus Grona Loureiro. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 11. | FNA vol. 11. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. wislizeni, Meibomia retinens | Hedysarum triflorum, Grona triflora, Meibomia triflora, Sagotia triflora |
Name authority | Schlechtendal: Linnaea 12: 311. (1838) | (Linnaeus) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 334. (1825) |
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