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Santa Rita Mountain ticktrefoil

Maryland tickclover, smooth small-leaf tick-trefoil

Habit Herbs, perennial, diffuse; base woody, rootstock woody. Herbs, perennial.
Stems

erect, ascending, or procumbent, usually striate, 30–60(–100) cm, sparsely to densely pilose and uncinate-pubescent.

ascending to erect, usually striate, branched or unbranched, 30–150 cm, glabrous or sparsely to densely uncinate-puberulent, sometimes also patent long-pilose.

Leaves

trifoliolate, sometimes unifoliolate proximally;

stipules persistent, recurved, subulate, 2–3 mm;

petiole 7–25 mm;

leaflet blades usually oblong-ovate to narrowly so, sometimes broadly elliptic to oblong, apex obtuse or acute, surfaces sparsely uncinate-puberulent;

terminal blade 10–30(–35) × 8–10 mm, length 1–3.5 times width.

trifoliolate;

stipules caducous or moderately persistent, subulate to narrowly ovate, 2–5 mm;

petiole 1–30 mm;

leaflet blades narrowly ovate, elliptic, elliptic-ovate, ovate-rhombic, or suborbiculate, apex obtuse or acute, mucronulate, surfaces sparsely to densely uncinate-puberulent and appressed-villous abaxially, uncinate-puberulent or glabrescent adaxially;

terminal blade 9–75 × 6–33 mm, length 1.5–4(–5) times width.

Inflorescences

usually unbranched, sometimes branched;

rachis patent uncinate-puberulent;

primary bracts narrowly ovate, 2.5–3 mm.

terminal and branched, axillary and unbranched;

rachis uncinate-puberulent;

primary bracts ovate, 1.5–3 mm.

Pedicels

8–15 mm.

3–19 mm, uncinate-puberulent.

Flowers

calyx 1.5–2.5 mm, pubescent, tube 1 mm;

abaxial lobes 1.5 mm, lateral lobes 1 mm;

corolla pink-purple, 4–5 mm.

calyx 1.5–2.5 mm, pubescent (not uncinate-puberulent), tube 1 mm;

abaxial lobes 1.2–1.5 mm, lateral lobes 1 mm;

corolla usually lavender to red-violet or pink-purple, rarely white, 4–6 mm.

Loments

sutures subequally crenate;

connections central, 1/5–1/4 as broad as segments;

segments 2–7, rounded or broadly elliptic, 2.5–3 × 2.5 mm, abaxially and adaxially rounded, glabrous or slightly puberulent;

stipe 1–2 mm.

sutures crenate abaxially, sinuate adaxially;

connections adaxial, 1/4–1/2 as broad as segments;

segments 1–4, broadly elliptic, 3.5–5.5 × 2.5–4 mm, rounded abaxially, convex adaxially, uncinate-puberulent throughout;

stipe 1–2 mm.

2n

= 22.

Desmodium retinens

Desmodium marilandicum

Phenology Flowering late summer–fall.
Habitat Wooded or grassy rocky slopes, oak or oak-pine forests.
Elevation 1400–2100 m. (4600–6900 ft.)
Distribution
from FNA
AZ; Mexico; Central America (Guatemala)
[BONAP county map]
from USDA
e United States; sc United States; West Indies
[WildflowerSearch map]
[BONAP county map]
Discussion

Desmodium retinens is known in the flora area from Cochise, Graham, Pima, and Santa Cruz counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Desmodium ciliare, D. lancifolium, and D. marilandicum have generally been recognized as closely related but distinct species in floras of the United States. They commonly have two or three articulate, small loments. D. Isely (1990, 1998) grouped them as the D. ciliare Group and regarded forms intermediate between them as putative hybrids: D. ciliare × D. marilandicum and D. ciliare × D. lancifolium. They have been separated by differences in the leaflets and pubescence, but the differences are not always clear. They are treated here as one species with three varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Terminal leaflet blades elliptic-ovate to narrowly ovate, 50–75 × 20–33 mm, apex acute to obtuse.
var. lancifolium
1. Terminal leaflet blades elliptic, ovate, ovate-rhombic, or suborbiculate, 9–40 × 6–17 mm, apex obtuse.
→ 2
2. Petioles 12–30 mm, sparsely uncinate-puberulent; stems with a few, scattered hairs; pedicels (6–)8–19 mm.
var. marilandicum
2. Petioles 1–15 mm; stems and petioles uncinate-puberulent and patent long-pilose; pedicels.
var. ciliare
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Desmodium Fabaceae > subfam. Faboideae > Desmodium
Sibling taxa
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
Subordinate taxa
D. marilandicum var. ciliare, D. marilandicum var. lancifolium, D. marilandicum var. marilandicum
Synonyms D. wislizeni, Meibomia retinens Hedysarum marilandicum, Meibomia marilandica
Name authority Schlechtendal: Linnaea 12: 311. (1838) (Linnaeus) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 328. (1825) — (as marylandicum)
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