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Santa Rita Mountain ticktrefoil

Habit Herbs, perennial, diffuse; base woody, rootstock woody. Herbs, perennial.
Stems

erect, ascending, or procumbent, usually striate, 30–60(–100) cm, sparsely to densely pilose and uncinate-pubescent.

prostrate, 100–200(–300) cm, sparsely to densely patent-pubescent and uncinate-puberulent.

Leaves

trifoliolate, sometimes unifoliolate proximally;

stipules persistent, recurved, subulate, 2–3 mm;

petiole 7–25 mm;

leaflet blades usually oblong-ovate to narrowly so, sometimes broadly elliptic to oblong, apex obtuse or acute, surfaces sparsely uncinate-puberulent;

terminal blade 10–30(–35) × 8–10 mm, length 1–3.5 times width.

trifoliolate;

stipules often caducous, sometimes persistent, ovate or narrowly ovate, 4.5–8 mm;

petiole 28–50 mm;

leaflet blades ovate or rhombic-ovate, lateral veins inconspicuous, arcuate along margin, apex acute or obtuse, surfaces sparsely strigose;

terminal blade 30–70 × 20–50 mm, length 1.4–2 times width.

Inflorescences

usually unbranched, sometimes branched;

rachis patent uncinate-puberulent;

primary bracts narrowly ovate, 2.5–3 mm.

lax-flowered, ascending, terminal and branched, also axillary and unbranched;

rachis uncinate-puberulent;

primary bracts caducous, ovate, 3–4 mm.

Pedicels

8–15 mm.

7–9 mm.

Flowers

calyx 1.5–2.5 mm, pubescent, tube 1 mm;

abaxial lobes 1.5 mm, lateral lobes 1 mm;

corolla pink-purple, 4–5 mm.

calyx 2.5–3.5 mm, uncinate-puberulent, tube 1 mm;

abaxial lobes 2 mm, lateral lobes 1.5 mm;

corolla purple, 8–9.5 mm.

Loments

sutures subequally crenate;

connections central, 1/5–1/4 as broad as segments;

segments 2–7, rounded or broadly elliptic, 2.5–3 × 2.5 mm, abaxially and adaxially rounded, glabrous or slightly puberulent;

stipe 1–2 mm.

sutures obtusely dentate abaxially, sinuate adaxially;

connections adaxial, 1/4–1/3 as broad as segments;

segments 3 or 4, deltate-rhombic, 6–8 × 4–5 mm, obtusely angled abaxially, straight or slightly convex adaxially, uncinate-pubescent;

stipe 2 mm.

Desmodium retinens

Desmodium ×humifusum

Phenology Flowering late summer–fall. Flowering summer–fall.
Habitat Wooded or grassy rocky slopes, oak or oak-pine forests. Woodland openings and edges, powerline cuts, near exposed limestone.
Elevation 1400–2100 m. (4600–6900 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
AZ; Mexico; Central America (Guatemala)
[BONAP county map]
from FNA
CT; DE; IN; MA; MD; MO; NJ; NY; PA
[BONAP county map]
Discussion

Desmodium retinens is known in the flora area from Cochise, Graham, Pima, and Santa Cruz counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Desmodium ×humifusum had been recognized as a species, but J. A. Raveill (2002) confirmed a hybrid origin from D. paniculatum and D. rotundifolium by allozyme electrophoresis. Desmodium × humifusum is likely to occur more widely than reported. It is in the Center for Plant Conservation’s National Collection of Endangered Plants as D. humifusum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Desmodium Fabaceae > subfam. Faboideae > Desmodium
Sibling taxa
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum
Synonyms D. wislizeni, Meibomia retinens Hedysaum humifusum, Meibomia humifusa
Name authority Schlechtendal: Linnaea 12: 311. (1838) (Muhlenberg ex Bigelow) L. C. Beck: Bot. North. Middle States, 86. (1833) — (as species)
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