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simple-leaf tick trefoil

arrow-leaf tick-trefoil, panicle tick-trefoil, panicle tickclover, panicledleaf ticktrefoil

Habit Herbs, perennial. Herbs, perennial; base woody, rootstock thick.
Stems

ascending to erect, branched, 30–80 cm, inconspicuously uncinate-puberulent or glabrescent.

erect or ascending, usually striate, 30–100(–150) cm, glabrous or sparsely to densely uncinate-puberulent or conspicuously pilose or glabrescent.

Leaves

unifoliolate;

stipules ± persistent, narrowly ovate-deltate, 3–3.5 mm;

petiole 6–11 mm;

leaflet blades ovate to narrowly ovate, 30–80 × 10–30 mm, length (2–)2.5–5 times width, apex acute or obtuse, with paler patches along midrib adaxially, surfaces uncinate-puberulent and villous.

trifoliolate;

stipules caducous, subulate to narrowly ovate-deltate, 2–6 mm;

petiole (10–)20–50 mm;

leaflet blades polymorphic, linear, narrowly ovate, narrowly ovate-oblong, ovate, broadly ovate to subrounded, or rhombic, usually thin, rarely thick, papery, apex acute to obtuse, surfaces closely to subdensely appressed pilose, inconspicuously reticulate-veined abaxially, sparsely appressed-puberulent and pilose adaxially;

terminal blade 20–100 × 8–65 mm, length 1–8 times width.

Inflorescences

often numerous, slender and flexuous, branched or unbranched;

rachis densely uncinate-puberulent and villous;

primary bracts narrowly ovate, 1–2 mm.

terminal panicles and axillary racemes, branched, very diffuse;

rachis moderately to densely uncinate-puberulent to -pubescent;

primary bracts narrowly ovate, 1–3.5 mm.

Pedicels

4–10 mm.

(3–)6–12(–20) mm.

Flowers

calyx 2.5–3 mm, puberulent and sparsely pilose, tube 1 mm;

abaxial lobes 2–3 mm, lateral lobes 2 mm, adaxial connate nearly to apex;

corolla pink to purple, 4–5 mm.

calyx 2–3 mm, pubescent, tube 1–1.5 mm;

abaxial lobes 2 mm, lateral lobes 1 mm;

corolla lilac to purple, 6–9 mm.

Loments

margins slightly involute when young, sutures crenate abaxially, sinuate adaxially;

connections adaxial, 1/5 as broad as segments;

segments 3–5, elliptic to obovate, 2.5–6 × 3–3.5 mm, rounded abaxially, convex adaxially, sparsely uncinate-puberulent, sutures glabrous;

stipe 1.5–2.5 mm.

sutures deeply crenate or dentate abaxially, sinuate adaxially;

connections adaxial, 1/5–1/3 as broad as segments;

segments (2 or)3–5, deltate to rhombic, (4–)5–10 × 3–5 mm, angled abaxially (sometimes obtusely), convex adaxially, uncinate-puberulent to -pubescent;

stipe 1–7 mm.

2n

= 22.

= 22.

Desmodium psilophyllum

Desmodium paniculatum

Phenology Flowering summer–fall.
Habitat Mountain woodlands, creek beds, terraces.
Elevation 1000–2200 m. (3300–7200 ft.)
Distribution
from FNA
AZ; TX; Mexico (Baja California Sur, Chiapas, Chihuahua, Coahuila, Durango, Hidalgo, Puebla, San Luis Potosí, Sinaloa, Sonora, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from USDA
North America [Introduced in e Asia (Japan)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Desmodium psilophyllum is known in the flora area from southern Arizona to western and central Texas at the edge of Edwards Plateau; it is not known from New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Desmodium paniculatum is allied with D. fernaldii, D. glabellum, and D. perplexum, and the four are called the D. paniculatum Group, which is characterized by having straight loments with 3–5 angled segments. These species have been studied intensively by B. G. Schubert (1950, 1950b) and D. Isely (1953, 1983b, 1990, 1998). According to Isely (1990, 1998), D. paniculatum intergrades with both D. perplexum and D. glabellum, resulting in a continuum of variation among the members of the D. paniculatum Group that suggests introgression, including: D. paniculatum × D. perplexum; D. paniculatum × D. glabellum; and D. glabellum × D. perplexum. They are treated here as varieties of a single polymorphic species. The varieties are distinguished by somewhat continuous or overlapping characters.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet blade surfaces slightly strigose to conspicuously subappressed-villous, sometimes uncinate-pubescent abaxially; stems and petioles glabrescent to conspicuously pilose or uncinate-pubescent.
var. paniculatum
1. Leaflet blade surfaces uncinate-puberulent on veins, sparsely strigulose abaxially; stems and petioles glabrous or uncinate-pubescent.
var. fernaldii
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Desmodium Fabaceae > subfam. Faboideae > Desmodium
Sibling taxa
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
Subordinate taxa
D. paniculatum var. fernaldii, D. paniculatum var. paniculatum
Synonyms Meibomia psilophylla Hedysarum paniculatum, Meibomia paniculata
Name authority Schlechtendal: Linnaea 12: 310. (1838) (Linnaeus) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 329. (1825)
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