The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Santa Cruz Island ticktrefoil

arrow-leaf tick-trefoil, panicle tick-trefoil, panicle tickclover, panicledleaf ticktrefoil

Habit Herbs or shrubs, perennial. Herbs, perennial; base woody, rootstock thick.
Stems

erect or ascending, branched, 40–100 cm, slightly uncinate-pubescent and pilose or glabrescent.

erect or ascending, usually striate, 30–100(–150) cm, glabrous or sparsely to densely uncinate-puberulent or conspicuously pilose or glabrescent.

Leaves

trifoliolate;

stipules mostly persistent, broadly ovate, 4–10 mm, base subamplexicaul;

petiole 30–40 mm;

leaflet blades broadly ovate-elliptic or ovate to narrowly ovate, apex acute, surfaces uncinate-puberulent;

terminal blade 20–70(–100) × 10–20(–50) mm, length 1.8–2 times width.

trifoliolate;

stipules caducous, subulate to narrowly ovate-deltate, 2–6 mm;

petiole (10–)20–50 mm;

leaflet blades polymorphic, linear, narrowly ovate, narrowly ovate-oblong, ovate, broadly ovate to subrounded, or rhombic, usually thin, rarely thick, papery, apex acute to obtuse, surfaces closely to subdensely appressed pilose, inconspicuously reticulate-veined abaxially, sparsely appressed-puberulent and pilose adaxially;

terminal blade 20–100 × 8–65 mm, length 1–8 times width.

Inflorescences

branched or unbranched;

rachis bulbous-villous and uncinate-puberulent;

primary bracts caducous, subulate, 1 mm.

terminal panicles and axillary racemes, branched, very diffuse;

rachis moderately to densely uncinate-puberulent to -pubescent;

primary bracts narrowly ovate, 1–3.5 mm.

Pedicels

10–20 mm, patent uncinate-pubescent.

(3–)6–12(–20) mm.

Flowers

calyx 1.5–2 mm, pilose and uncinate-puberulent, tube 0.8 mm;

abaxial lobes 1 mm, lateral lobes 0.8–1 mm;

corolla pink-purple, fading greenish, 4–5 mm.

calyx 2–3 mm, pubescent, tube 1–1.5 mm;

abaxial lobes 2 mm, lateral lobes 1 mm;

corolla lilac to purple, 6–9 mm.

Loments

sutures subequally deeply crenate;

connections central, 1/8–1/7 as broad as segments;

segments 3–6, rounded, 6–10 × 5–7 mm, rounded, sparsely pubescent, at least on sutures;

stipe 1(–2) mm.

sutures deeply crenate or dentate abaxially, sinuate adaxially;

connections adaxial, 1/5–1/3 as broad as segments;

segments (2 or)3–5, deltate to rhombic, (4–)5–10 × 3–5 mm, angled abaxially (sometimes obtusely), convex adaxially, uncinate-puberulent to -pubescent;

stipe 1–7 mm.

2n

= 22.

Desmodium psilocarpum

Desmodium paniculatum

Phenology Flowering late summer–fall.
Habitat Woodland, grasslands, canyon slopes, streamsides.
Elevation 1000–2000 m. (3300–6600 ft.)
Distribution
from FNA
AZ; NM; Mexico (Chihuahua, Nuevo León, Sonora, Tamaulipas)
[BONAP county map]
from USDA
North America [Introduced in e Asia (Japan)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Desmodium psilocarpum is known in the flora area from southern Arizona and adjacent southwestern New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Desmodium paniculatum is allied with D. fernaldii, D. glabellum, and D. perplexum, and the four are called the D. paniculatum Group, which is characterized by having straight loments with 3–5 angled segments. These species have been studied intensively by B. G. Schubert (1950, 1950b) and D. Isely (1953, 1983b, 1990, 1998). According to Isely (1990, 1998), D. paniculatum intergrades with both D. perplexum and D. glabellum, resulting in a continuum of variation among the members of the D. paniculatum Group that suggests introgression, including: D. paniculatum × D. perplexum; D. paniculatum × D. glabellum; and D. glabellum × D. perplexum. They are treated here as varieties of a single polymorphic species. The varieties are distinguished by somewhat continuous or overlapping characters.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet blade surfaces slightly strigose to conspicuously subappressed-villous, sometimes uncinate-pubescent abaxially; stems and petioles glabrescent to conspicuously pilose or uncinate-pubescent.
var. paniculatum
1. Leaflet blade surfaces uncinate-puberulent on veins, sparsely strigulose abaxially; stems and petioles glabrous or uncinate-pubescent.
var. fernaldii
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Desmodium Fabaceae > subfam. Faboideae > Desmodium
Sibling taxa
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
Subordinate taxa
D. paniculatum var. fernaldii, D. paniculatum var. paniculatum
Synonyms Meibomia psilocarpa Hedysarum paniculatum, Meibomia paniculata
Name authority A. Gray: Smithsonian Contr. Knowl. 5(6): 48. (1853) (Linnaeus) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 329. (1825)
Web links