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Maryland tickclover, smooth small-leaf tick-trefoil

pine barren ticktrefoil

Habit Herbs, perennial. Herbs, perennial.
Stems

ascending to erect, usually striate, branched or unbranched, 30–150 cm, glabrous or sparsely to densely uncinate-puberulent, sometimes also patent long-pilose.

ascending or erect, usually striate, unbranched to inflorescence, 50–100 cm, uncinate-puberulent.

Leaves

trifoliolate;

stipules caducous or moderately persistent, subulate to narrowly ovate, 2–5 mm;

petiole 1–30 mm;

leaflet blades narrowly ovate, elliptic, elliptic-ovate, ovate-rhombic, or suborbiculate, apex obtuse or acute, mucronulate, surfaces sparsely to densely uncinate-puberulent and appressed-villous abaxially, uncinate-puberulent or glabrescent adaxially;

terminal blade 9–75 × 6–33 mm, length 1.5–4(–5) times width.

trifoliolate;

stipules mostly persistent, linear to narrowly ovate-deltate, 2–5 mm;

petiole 6.5–18 mm;

leaflet blades linear to narrowly oblong, usually folded and appearing narrower, leathery, apex obtuse to acute, surfaces reticulate-veined adaxially, glabrous or sparsely puberulent abaxially, glabrous adaxially;

terminal blade 35–60(–80) × 4–7 mm, length 8–10 times width.

Inflorescences

terminal and branched, axillary and unbranched;

rachis uncinate-puberulent;

primary bracts ovate, 1.5–3 mm.

branched or unbranched;

rachis densely uncinate-puberulent to pubescent;

primary bracts narrowly lanceolate-triangular, 1.4–2 mm.

Pedicels

3–19 mm, uncinate-puberulent.

5–13 mm.

Flowers

calyx 1.5–2.5 mm, pubescent (not uncinate-puberulent), tube 1 mm;

abaxial lobes 1.2–1.5 mm, lateral lobes 1 mm;

corolla usually lavender to red-violet or pink-purple, rarely white, 4–6 mm.

calyx 2.5–3.5 mm, uncinate-puberulent to pubescent on lobes, tube 1 mm;

abaxial lobes 2–2.5 mm, lateral lobes 1.5–2 mm;

corolla pink or purple, 4 mm.

Loments

sutures crenate abaxially, sinuate adaxially;

connections adaxial, 1/4–1/2 as broad as segments;

segments 1–4, broadly elliptic, 3.5–5.5 × 2.5–4 mm, rounded abaxially, convex adaxially, uncinate-puberulent throughout;

stipe 1–2 mm.

sutures deeply crenate abaxially, with narrow connection between segments, nearly straight adaxially, slightly sinuate at isthmus;

connections adaxial, 1/5 as broad as segments;

segments 1 or 2(or 3), semiorbiculate, 4.5–6 × 3–4 mm, symmetrically rounded abaxially, incipiently or plainly concave adaxially, densely uncinate-puberulent, more densely on sutures;

stipe 1.5 mm.

2n

= 22.

= 22.

Desmodium marilandicum

Desmodium strictum

Phenology Flowering late summer–fall.
Habitat Dry woodland, barren sandhills, ruderal areas.
Elevation 0–500 m. (0–1600 ft.)
Distribution
from USDA
e United States; sc United States; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; DC; DE; FL; GA; LA; MD; MO; MS; NC; NJ; OK; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Desmodium ciliare, D. lancifolium, and D. marilandicum have generally been recognized as closely related but distinct species in floras of the United States. They commonly have two or three articulate, small loments. D. Isely (1990, 1998) grouped them as the D. ciliare Group and regarded forms intermediate between them as putative hybrids: D. ciliare × D. marilandicum and D. ciliare × D. lancifolium. They have been separated by differences in the leaflets and pubescence, but the differences are not always clear. They are treated here as one species with three varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Desmodium strictum is mostly restricted to dry, sandy soils along the Atlantic and Gulf coasts. Because of the close similarities between D. strictum and D. tenuifolium, mature fruits are necessary to distinguish the two species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Terminal leaflet blades elliptic-ovate to narrowly ovate, 50–75 × 20–33 mm, apex acute to obtuse.
var. lancifolium
1. Terminal leaflet blades elliptic, ovate, ovate-rhombic, or suborbiculate, 9–40 × 6–17 mm, apex obtuse.
→ 2
2. Petioles 12–30 mm, sparsely uncinate-puberulent; stems with a few, scattered hairs; pedicels (6–)8–19 mm.
var. marilandicum
2. Petioles 1–15 mm; stems and petioles uncinate-puberulent and patent long-pilose; pedicels.
var. ciliare
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Desmodium Fabaceae > subfam. Faboideae > Desmodium
Sibling taxa
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
Subordinate taxa
D. marilandicum var. ciliare, D. marilandicum var. lancifolium, D. marilandicum var. marilandicum
Synonyms Hedysarum marilandicum, Meibomia marilandica Hedysarum strictum, Meibomia stricta
Name authority (Linnaeus) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 328. (1825) — (as marylandicum) (Pursh) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 329. (1825)
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