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Lindheimer's ticktrefoil

creeping beggarweed, Spanish clover, Spanish tick-trefoil, zarzabacoa comun

Habit Herbs, perennial. Herbs, subshrubs, or shrubs, perennial; stoloniferous or rhizomatous.
Stems

erect, branched, angled and grooved, 40–60 cm, uncinate-puberulent and -pubescent, also sparsely pilose.

erect or ascending, to 300 cm, pubescent or glabrescent.

Leaves

trifoliolate;

stipules ± persistent, reflexed, broadly or obliquely ovate, 7–8 mm, base subamplexicaul;

petiole 10–50 mm;

leaflet blades usually ovate, rhombic to broadly ovate, or semiorbiculate, rarely narrowly ovate, base truncate or broadly obtuse, apex acute, surfaces densely villosulous abaxially, uncinate-puberulent and pilose adaxially;

terminal blade 50–100 × 40–75 mm, length usually less than 2 times width.

trifoliolate;

stipules usually persistent, narrowly ovate-deltate, 5–10 mm;

petiole usually 15–20 mm;

leaflet blades elliptic to ovate, apex obtuse or acute, surfaces finely spreading-villosulous to substrigose abaxially, uncinate-puberulent or glabrescent adaxially;

terminal blade 20–90 × 15–45 mm, length 1.5–4 times width.

Inflorescences

usually unbranched;

rachis densely patent uncinate-pubescent;

primary bracts broadly ovate, 4.5–7 mm.

unbranched;

rachis densely patent uncinate-pubescent;

primary bracts caducous, narrowly ovate, 6–7 mm.

Pedicels

5 mm.

persistent with calyx-remnant at top after loments drop, 5–9 mm.

Flowers

calyx 2–2.5 mm, uncinate-puberulent, tube 1 mm;

abaxial lobes 1–1.5 mm, lateral lobes 0.7 mm;

corolla blue-green, 6–7 mm.

calyx 2–3.5 mm, uncinate-puberulent, lobes pilose, tube 1 mm;

abaxial lobes 1.5–2.5 mm, lateral lobes 1–2 mm;

corolla purple, 5–8 mm.

Loments

margins ± involute and contorted, sutures deeply crenate abaxially, shallowly dentate adaxially;

connections central, 1/5 as broad as segments;

segments (1–)3–5, subrhombic to semiovate, 7–15 × 5–8 mm, rounded abaxially, obtusely angled adaxially, glabrous, sutures densely uncinate-pubescent;

stipe 2–3.5 mm.

sutures symmetrically crenate abaxially, straight or slightly sinuate adaxially;

connections central, 1/2–2/3 as broad as segments;

segments 4–8, semiobovate, 3.5–5 × 2.5–3 mm, broadly rounded abaxially, straight or barely convex adaxially, uncinate-puberulent;

stipe 1.5–2 mm.

2n

= 22.

Desmodium lindheimeri

Desmodium incanum

Phenology Flowering summer–fall. Flowering spring–fall.
Habitat Rocky ravines, oak-juniper woodlands. Pine-palmetto flatwoods, woodland borders, lawns, ruderal sites, disturbed or waste areas.
Elevation 200–300 m. (700–1000 ft.) 0–50 m. (0–200 ft.)
Distribution
from FNA
TX; Mexico (Chiapas, Coahuila, Durango, Nuevo León, San Luis Potosí, Tamaulipas)
[BONAP county map]
from FNA
AL; FL; GA; TX; Central America; South America; Mexico (Chiapas, Jalisco, Oaxaca, Quintana Roo, Tamaulipas, Veracruz); West Indies [Introduced in North America; introduced also in Asia (Taiwan), Africa, Indian Ocean Islands (Mauritius, Reunion), Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Desmodium lindheimeri resembles D. canescens and its relatives, especially D. ochroleucum, in having relatively large loments. It is common in its narrow range in the Edwards Plateau (Comal County) and is widespread in Mexico (M. Enquist 1995).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Desmodium incanum may be distinguished by its long-persistent stipules usually fused and nearly surrounding the stem, at least when young, and by its pedicels which are usually borne singly and are each subtended by one primary bract and two (lateral) secondary bracts (B. G. Schubert 1980).

Desmodium incanum was long known as D. canum Schinz & Thellung (= Meibomia cana S. F. Blake) based on the illegitimate Hedysarum canum J. F. Gmelin, a superfluous name for H. racemosum Aublet. The complex nomenclatural history was elaborated by D. H. Nicolson (1978) and L. C. P. Lima et al. (2012, 2014). Hedysarum canescens Miller (1768) is a later homonym of H. canescens Linnaeus (1753), thus illegitimate, and pertains here. Hedysarum canum J. F. Gmelin is a superfluous name for H. racemosum Aublet; Meibomia cana S. F. Blake was intended as a new combination based on that name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Desmodium Fabaceae > subfam. Faboideae > Desmodium
Sibling taxa
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. incanum, D. intortum, D. laevigatum, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
D. arizonicum, D. batocaulon, D. canadense, D. canescens, D. cinerascens, D. cuspidatum, D. floridanum, D. grahamii, D. gramineum, D. illinoense, D. intortum, D. laevigatum, D. lindheimeri, D. lineatum, D. marilandicum, D. metcalfei, D. nuttallii, D. ochroleucum, D. paniculatum, D. procumbens, D. psilocarpum, D. psilophyllum, D. retinens, D. rosei, D. rotundifolium, D. scopulorum, D. scorpiurus, D. sessilifolium, D. strictum, D. tenuifolium, D. tortuosum, D. triflorum, D. tweedyi, D. viridiflorum, D. ×humifusum
Synonyms Meibomia lindheimeri Hedysarum incanum, Aeschynomene incana, D. ancistrocarpum, D. canum, D. frutescens, D. frutescens var. amplyophyllum, D. malacophyllum, D. mauritianum, D. supinum var. amblyophyllum, H. ancistrocarpum, H. malacophyllum, H. mauritianum, H. racemosum, Meibomia adscendens var. incana, M. incana, M. malacophylla, M. supina
Name authority Vail: Bull. Torrey Bot. Club 18: 120. (1891) (Swartz) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 332. (1825) — name conserved
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