Desmodium intortum |
Desmodium marilandicum |
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greenleaf ticktrefoil |
Maryland tickclover, smooth small-leaf tick-trefoil |
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Habit | Herbs, perennial. | Herbs, perennial. | ||||||||
Stems | prostrate or climbing, branched, angular (3-sided), 40–200 cm, uncinate-pubescent and villous. |
ascending to erect, usually striate, branched or unbranched, 30–150 cm, glabrous or sparsely to densely uncinate-puberulent, sometimes also patent long-pilose. |
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Leaves | trifoliolate; stipules usually persistent, sometimes caducous, ovate-deltate, 5.5–12 mm; petiole 4–8 cm; leaflet blades ovate, elliptic to broadly ovate, or rhombic, apex acute to acuminate, surfaces more densely sericeous abaxially, sparsely to densely sericeous adaxially; terminal blade 35–100 × 24–60 mm, length 2.5–3 times width. |
trifoliolate; stipules caducous or moderately persistent, subulate to narrowly ovate, 2–5 mm; petiole 1–30 mm; leaflet blades narrowly ovate, elliptic, elliptic-ovate, ovate-rhombic, or suborbiculate, apex obtuse or acute, mucronulate, surfaces sparsely to densely uncinate-puberulent and appressed-villous abaxially, uncinate-puberulent or glabrescent adaxially; terminal blade 9–75 × 6–33 mm, length 1.5–4(–5) times width. |
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Inflorescences | branched or unbranched; rachis patent-villous and uncinate-pubescent to puberulent; primary bracts densely imbricate before anthesis, caducous, narrowly ovate, 7–10 mm. |
terminal and branched, axillary and unbranched; rachis uncinate-puberulent; primary bracts ovate, 1.5–3 mm. |
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Pedicels | 4–8 mm. |
3–19 mm, uncinate-puberulent. |
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Flowers | calyx 3–5 mm, sparsely puberulent throughout, lobes often villous, tube 1 mm; abaxial lobes 4–5.5 mm, lateral lobes 2.5–4 mm; corolla purple, fading blue-green, 7–9 mm. |
calyx 1.5–2.5 mm, pubescent (not uncinate-puberulent), tube 1 mm; abaxial lobes 1.2–1.5 mm, lateral lobes 1 mm; corolla usually lavender to red-violet or pink-purple, rarely white, 4–6 mm. |
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Loments | sutures crenate or somewhat angled abaxially, sinuate adaxially; connections adaxial, 1/2 as broad as segments; segments 5–9, semiorbiculate, 3–4 × 2–2.5 mm, obliquely rounded abaxially, slightly angled adaxially, uncinate-pubescent; stipe 1–2 mm. |
sutures crenate abaxially, sinuate adaxially; connections adaxial, 1/4–1/2 as broad as segments; segments 1–4, broadly elliptic, 3.5–5.5 × 2.5–4 mm, rounded abaxially, convex adaxially, uncinate-puberulent throughout; stipe 1–2 mm. |
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2n | = 22. |
= 22. |
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Desmodium intortum |
Desmodium marilandicum |
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Phenology | Flowering late summer–fall. | |||||||||
Habitat | Open, grassy areas, roadsides. | |||||||||
Elevation | 1000–1700 m. (3300–5600 ft.) | |||||||||
Distribution |
AZ; Central America; Mexico (Chiapas, Colima, Guerrero, Michoacán, Sonora, Veracruz) [Introduced in Pacific Islands (Hawaii), Australia] |
e United States; sc United States; West Indies
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Discussion | In the flora area, Desmodium intortum is known only from Cochise County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Desmodium ciliare, D. lancifolium, and D. marilandicum have generally been recognized as closely related but distinct species in floras of the United States. They commonly have two or three articulate, small loments. D. Isely (1990, 1998) grouped them as the D. ciliare Group and regarded forms intermediate between them as putative hybrids: D. ciliare × D. marilandicum and D. ciliare × D. lancifolium. They have been separated by differences in the leaflets and pubescence, but the differences are not always clear. They are treated here as one species with three varieties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 11. | FNA vol. 11. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Hedysarum intortum, D. sonorae, D. uncinatum, Meibomia intorta | Hedysarum marilandicum, Meibomia marilandica | ||||||||
Name authority | (Miller) Urban: Symb. Antill. 8: 292. (1920) | (Linnaeus) de Candolle in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 2: 328. (1825) — (as marylandicum) | ||||||||
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