Descurainia sophioides |
Descurainia paradisa |
|
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northern tansy-mustard |
Great Basin tansy mustard, Nevada tansymustard, paradise tansymustard |
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Habit | Annuals or biennials; eglandular or glandular distally; glabrate to moderately pubescent, trichomes dendritic, sometimes mixed with simple ones. | Annuals; glandular or eglandular; sparsely to densely pubescent, trichomes dendritic. |
Stems | erect, unbranched or sometimes branched distally, (0.5–)1.5–11(–18) dm. |
erect, branched basally and distally, (often purplish), (1–)1.5–3.2(–4.1) dm. |
Basal leaves | petiole 0.5–5 cm; blade pinnate or, sometimes, 2-pinnate, broadly oblanceolate to ovate in outline, 2.5–11.4(–15.2) cm, lateral lobes lanceolate, (to 10 × 4 mm), margins incised. |
(soon withered); petiole 0.3–1.3 cm; blade pinnate, oblanceolate to obovate in outline, 1.5–3 cm, lateral lobes oblong to linear or lanceolate, (1–5 × 0.3–1 mm), margins entire or dentate. |
Cauline leaves | sessile or shortly petiolate; blade smaller distally, distal lobes often narrower, surfaces often glabrous or sparsely pubescent. |
sessile; blade smaller distally, distal lobes often narrower, surfaces moderately to densely pubescent. |
Racemes | elongated or not in fruit, (flowers overtopped by developing fruits). |
considerably elongated in fruit. |
Flowers | sepals erect, yellowish, oblong, 1.6–2.7 mm, glabrous; petals narrowly oblanceolate, 2–2.5 × 0.3–0.6 mm; median filaments 2.5–3.5 mm; anthers 0.3–0.4 mm. |
sepals spreading to ascending, pale yellow, oblong, 0.8–1.2 mm, pubescent; petals oblanceolate, 0.9–1.3 × 0.2–0.5 mm; median filaments 0.8–1.2 mm; anthers 0.1–0.2 mm. |
Fruiting pedicels | divaricate to ascending, (often recurved in age), slender, (3–)4–9(–13) mm. |
divaricate to ascending, straight, 2.5–7(–9) mm. |
Fruits | erect to widely spreading, narrowly linear, slightly torulose, (9–)14–30(–34) × 0.6–1.1 mm, (usually terete, rarely slightly flattened, often curved inward); valves each with obscure midvein; septum not veined; ovules 30–62 per ovary; style obsolete, 0.07–0.3 mm, glabrous. |
divaricate to erect, usually obovoid to clavate, rarely broadly ellipsoid, not torulose, 2–5 × 1–2 mm, (acute basally, obtuse apically); valves each with obscure midvein; septum not veined; ovules 4–10 per ovary; style 0.05–0.3 mm, glabrous. |
Seeds | uniseriate, light brown, narrowly oblong, 1–1.5 × 0.3–0.5 mm. |
uniseriate or biseriate, brown, oblong, 0.8–1.2 × 0.5–0.6 mm. |
2n | = 14. |
|
Descurainia sophioides |
Descurainia paradisa |
|
Phenology | Flowering Jun–Sep. | Flowering Apr–Jun. |
Habitat | Open meadows, eroded peat, roadsides, disturbed and waste sites, rocky outcrops, mining dumps, gravelly grounds, stream banks, gullies | Shrub communities, sandy washes and dunes, roadsides |
Elevation | 0-1000 m (0-3300 ft) | 1000-2300 m (3300-7500 ft) |
Distribution |
AK; BC; MB; NT; NU; YT; Asia (Russian Far East, Siberia) |
CA; NV; OR
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Discussion | Although L. E. Detling (1939) reduced Descurainia paradisa to a subspecies of D. pinnata, molecular data (B. E. Goodson 2007) clearly show that it should not be included in that species. The boundaries of D. paradisa in its northern and southern ranges tend to be blurred relative to D. nelsonii and D. pinnata, respectively. R. C. Rollins (1993) and N. H. Holmgren (2005b) recognized some of the Nevada plants of Descurainia paradisa that have eglandular racemes as a subspecies and variety (nevadensis), respectively. This poorly delimited division of the species is artificial, and both glandular and eglandular forms are sometimes found within individual populations of most species. They also indicated that nevadensis has styles 0.2–0.3 mm (versus 0.05–0.15 mm in D. paradisa), but this distinction is equally unreliable. Indeed, the style length and the presence versus absence of glands are not inherited together. Some of the eglandular plants have styles to 0.1 mm (e.g., Williams & Tiehm 86-51-1, GH). It is likely that some of the plants identified as nevadensis are of hybrid origin involving other species, especially D. nelsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 529. | FNA vol. 7, p. 526. |
Parent taxa | Brassicaceae > tribe Descurainieae > Descurainia | Brassicaceae > tribe Descurainieae > Descurainia |
Sibling taxa | ||
Synonyms | Sisymbrium sophioides, Hesperis arctica, Sisymbrium arcticum, Sophia sophioides | Sophia paradisa, D. paradisa subsp. nevadensis, D. paradisa var. nevadensis, D. pinnata subsp. paradisa, D. pinnata var. paradisa, Sisymbrium paradisum |
Name authority | (Fischer ex Hooker) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 86[IV,105]: 316. (1924) | (A. Nelson & P. B. Kennedy) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 86[IV,105]: 331. (1924) |
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