Descurainia sophioides |
Descurainia nelsonii |
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northern tansy-mustard |
Nelson's tansy-mustard, sagebrush tansymustard |
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Habit | Annuals or biennials; eglandular or glandular distally; glabrate to moderately pubescent, trichomes dendritic, sometimes mixed with simple ones. | Annuals; eglandular; sparsely to moderately pubescent, sometimes glabrous distally, not canescent, trichomes dendritic. |
Stems | erect, unbranched or sometimes branched distally, (0.5–)1.5–11(–18) dm. |
erect, usually branched basally or slightly distally, rarely unbranched, (0.7–)0.9–3.2(–4.5) dm. |
Basal leaves | petiole 0.5–5 cm; blade pinnate or, sometimes, 2-pinnate, broadly oblanceolate to ovate in outline, 2.5–11.4(–15.2) cm, lateral lobes lanceolate, (to 10 × 4 mm), margins incised. |
petiole 0.5–1.5 cm; blade pinnate, ovate or oblong in outline, 0.8–2.5 cm, lateral lobe (2–5 pairs), margins dentate or entire. |
Cauline leaves | sessile or shortly petiolate; blade smaller distally, distal lobes often narrower, surfaces often glabrous or sparsely pubescent. |
sessile or shortly petiolate; blade smaller distally, distal lobes often narrower, sparsely to moderately pubescent. |
Racemes | elongated or not in fruit, (flowers overtopped by developing fruits). |
considerably elongated in fruit. |
Flowers | sepals erect, yellowish, oblong, 1.6–2.7 mm, glabrous; petals narrowly oblanceolate, 2–2.5 × 0.3–0.6 mm; median filaments 2.5–3.5 mm; anthers 0.3–0.4 mm. |
sepals ascending, yellowish, oblong, 0.7–1.2 mm, pubescent; petals narrowly oblanceolate, 0.8–1.2 × 0.2–0.4 mm; median filaments 1–1.5 mm; anthers 0.1–0.2 mm. |
Fruiting pedicels | divaricate to ascending, (often recurved in age), slender, (3–)4–9(–13) mm. |
divaricate-ascending (often at 20–45º angle), straight, (1.5–)2.5–7(–10) mm. |
Fruits | erect to widely spreading, narrowly linear, slightly torulose, (9–)14–30(–34) × 0.6–1.1 mm, (usually terete, rarely slightly flattened, often curved inward); valves each with obscure midvein; septum not veined; ovules 30–62 per ovary; style obsolete, 0.07–0.3 mm, glabrous. |
erect or ascending, linear, not or slightly torulose, (0.4–)5–8(–10) × 0.7–1 mm; valves each with distinct midvein; septum not veined; ovules 6–12 per ovary; style 0.1–0.2 mm, glabrous. |
Seeds | uniseriate, light brown, narrowly oblong, 1–1.5 × 0.3–0.5 mm. |
uniseriate, brown, oblong, 0.6–0.8 × 0.4–0.5 mm. |
2n | = 14. |
= 14. |
Descurainia sophioides |
Descurainia nelsonii |
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Phenology | Flowering Jun–Sep. | Flowering late May-mid Jul. |
Habitat | Open meadows, eroded peat, roadsides, disturbed and waste sites, rocky outcrops, mining dumps, gravelly grounds, stream banks, gullies | Roadsides, sagebrush, wash bottoms, silty flats, gravelly grounds |
Elevation | 0-1000 m (0-3300 ft) | 800-3000 m (2600-9800 ft) |
Distribution |
AK; BC; MB; NT; NU; YT; Asia (Russian Far East, Siberia) |
CA; ID; MT; NV; OR; WA; WY; BC
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Discussion | Descurainia nelsonii was treated by L. E. Detling (1939) and R. C. Rollins (1993) as a subspecies of D. pinnata, but the latter in the sense of these authors is not monophyletic, comprising instead either four or two unrelated species, respectively. ITS molecular data (B. E. Goodson 2007) suggest that D. nelsonii is most closely related to D. longepedicellata and D. paradisa. It can be distinguished from the latter species by its linear fruits with cuneate tips; D. paradisa has obovoid fruits with rounded tips. Descurainia nelsonii resembles D. pinnata subsp. brachycarpa in the orientation of fruiting pedicels and in having short styles (to 0.3 mm) and small seeds (to 1 × 0.5 mm). It differs in being branched (versus simple) at base and in having smaller flowers (petals 0.7–1 versus 1.5–2.6 mm), fewer ovules (6–12 versus 16–40) per ovary, linear (versus subclavate) fruits, and uniseriate (versus biseriate) seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 529. | FNA vol. 7, p. 525. |
Parent taxa | Brassicaceae > tribe Descurainieae > Descurainia | Brassicaceae > tribe Descurainieae > Descurainia |
Sibling taxa | ||
Synonyms | Sisymbrium sophioides, Hesperis arctica, Sisymbrium arcticum, Sophia sophioides | Sophia nelsonii, D. brachycarpa var. nelsonii, D. pinnata subsp. nelsonii, D. pinnata var. nelsonii |
Name authority | (Fischer ex Hooker) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 86[IV,105]: 316. (1924) | (Rydberg) Al-Shehbaz & Goodson: Harvard Pap. Bot. 12: 422. (2007) |
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