Descurainia sophioides |
Descurainia longepedicellata |
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northern tansy-mustard |
mountain tansymustard, narrow tansymustard, sticky tansymustard, western tansy-mustard |
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Habit | Annuals or biennials; eglandular or glandular distally; glabrate to moderately pubescent, trichomes dendritic, sometimes mixed with simple ones. | Annuals; usually eglandular, rarely glandular; moderately to sparsely pubescent, often glabrous distally, not canescent, trichomes dendritic. |
Stems | erect, unbranched or sometimes branched distally, (0.5–)1.5–11(–18) dm. |
erect, unbranched or branched proximally, often branched distally, (1.5–)3–6.2(–8.5) dm. |
Basal leaves | petiole 0.5–5 cm; blade pinnate or, sometimes, 2-pinnate, broadly oblanceolate to ovate in outline, 2.5–11.4(–15.2) cm, lateral lobes lanceolate, (to 10 × 4 mm), margins incised. |
petiole 0.4–3.5 cm; blade pinnate, ovate to oblanceolate in outline, 1.5–7 cm, lateral lobes linear or oblong, margins entire or dentate to incised. |
Cauline leaves | sessile or shortly petiolate; blade smaller distally, distal lobes often narrower, surfaces often glabrous or sparsely pubescent. |
sessile or shortly petiolate; blade smaller distally, lobes linear to filiform, margins entire, surfaces usually glabrous, rarely pubescent. |
Racemes | elongated or not in fruit, (flowers overtopped by developing fruits). |
considerably elongated in fruit. |
Flowers | sepals erect, yellowish, oblong, 1.6–2.7 mm, glabrous; petals narrowly oblanceolate, 2–2.5 × 0.3–0.6 mm; median filaments 2.5–3.5 mm; anthers 0.3–0.4 mm. |
sepals ascending, yellow, oblong, 1.5–2 mm, glabrous; petals narrowly oblanceolate, 1.7–2.6 cm × 0.5–1 mm; median filaments 1.5–2 mm; anthers 0.3–0.4 mm. |
Fruiting pedicels | divaricate to ascending, (often recurved in age), slender, (3–)4–9(–13) mm. |
horizontal to divaricate, straight, (8–)10–15(–20) mm. |
Fruits | erect to widely spreading, narrowly linear, slightly torulose, (9–)14–30(–34) × 0.6–1.1 mm, (usually terete, rarely slightly flattened, often curved inward); valves each with obscure midvein; septum not veined; ovules 30–62 per ovary; style obsolete, 0.07–0.3 mm, glabrous. |
erect, linear, not torulose, (9–)12–17 × 0.8–1.1 mm, (straight or slightly curved inward); valves each with obscure midvein; septum not veined; ovules 18–32 per ovary; style 0.1–0.2 mm, glabrous. |
Seeds | uniseriate, light brown, narrowly oblong, 1–1.5 × 0.3–0.5 mm. |
uniseriate, reddish brown, oblong, 1–1.3 × 0.6–0.7 mm. |
2n | = 14. |
= 14. |
Descurainia sophioides |
Descurainia longepedicellata |
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Phenology | Flowering Jun–Sep. | Flowering Apr–Jul. |
Habitat | Open meadows, eroded peat, roadsides, disturbed and waste sites, rocky outcrops, mining dumps, gravelly grounds, stream banks, gullies | Sandy plains and banks, dry washes, open hillsides, sagebrush and juniper or pine communities, grasslands |
Elevation | 0-1000 m (0-3300 ft) | 200-2100 m (700-6900 ft) |
Distribution |
AK; BC; MB; NT; NU; YT; Asia (Russian Far East, Siberia) |
AZ; CO; ID; MT; NV; OR; UT; WA; WY; BC
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Discussion | L. E. Detling (1939) treated Descurainia longepedicellata as subsp. filipes of D. pinnata, whereas R. C. Rollins (1993) and N. H. Holmgren (2005b) treated it as a subspecies and variety, respectively, of D. incisa. Molecular data, both nuclear and plastidic (B. E. Goodson 2007), place the three taxa in different, well-supported clades. R. C. Rollins (1993) and N. H. Holmgren (2005b) reported 2n = 28 and 42 for Descurainia longepedicellata (as D. pinnata var. filipes), but these counts are not vouchered. Rollins indicated that the taxon range extends into California and New Mexico; we have not seen material from those states. Descurainia longepedicellata resembles D. incisa subsp. paysonii in having long fruiting pedicels and linear leaf lobes with entire margins. The latter is easily distinguished by being canescent (versus not canescent) and having fruits strongly curved inward (versus straight). Because the two taxa are not closely related (B. E. Goodson 2007), the similarities in fruiting pedicels and distalmost leaf segments represent convergence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 529. | FNA vol. 7, p. 524. |
Parent taxa | Brassicaceae > tribe Descurainieae > Descurainia | Brassicaceae > tribe Descurainieae > Descurainia |
Sibling taxa | ||
Synonyms | Sisymbrium sophioides, Hesperis arctica, Sisymbrium arcticum, Sophia sophioides | Sisymbrium longepedicellatum, D. brachycarpa var. eglandulosa, D. incisa subsp. filipes, D. incisa var. filipes, D. longepedicellata var. glandulosa, D. pinnata subsp. filipes, D. pinnata var. filipes, D. rydbergii var. eglandulosa, Hesperis longepedicellata, Sisymbrium brachycarpum var. filipes, Sisymbrium incisum var. filipes, Sisymbrium incisum var. xerophilum, Sisymbrium longepedicellatum var. glandulosum, Sophia filipes, Sophia glandifera, Sophia gracilis, Sophia longepedicellata |
Name authority | (Fischer ex Hooker) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 86[IV,105]: 316. (1924) | (E. Fournier) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 86[IV,105]: 324. (1924) |
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