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northern tansy-mustard

Habit Annuals or biennials; eglandular or glandular distally; glabrate to moderately pubescent, trichomes dendritic, sometimes mixed with simple ones. Biennials; eglandular; finely pubescent, often canescent, trichomes dendritic, rarely also mixed with simple ones.
Stems

erect, unbranched or sometimes branched distally, (0.5–)1.5–11(–18) dm.

erect, unbranched basally, branched distally, 6–11 dm.

Basal leaves

petiole 0.5–5 cm;

blade pinnate or, sometimes, 2-pinnate, broadly oblanceolate to ovate in outline, 2.5–11.4(–15.2) cm, lateral lobes lanceolate, (to 10 × 4 mm), margins incised.

petiole 0.5–1.5 cm;

blade pinnate, oblanceolate to obovate or ovate in outline, 1–5 cm, lateral lobes (2–5 pairs), linear, margins entire or serrate to incised (apex obtuse).

Cauline leaves

sessile or shortly petiolate;

blade smaller distally, distal lobes often narrower, surfaces often glabrous or sparsely pubescent.

sessile or shortly petiolate;

blade smaller distally, distal lobes often narrower, surfaces densely pubescent.

Racemes

elongated or not in fruit, (flowers overtopped by developing fruits).

(paniculate, often with short branches), considerably elongated in fruit.

Flowers

sepals erect, yellowish, oblong, 1.6–2.7 mm, glabrous;

petals narrowly oblanceolate, 2–2.5 × 0.3–0.6 mm;

median filaments 2.5–3.5 mm;

anthers 0.3–0.4 mm.

sepals spreading, yellowish, oblong, 1.4–2 mm, pubescent, (trichomes dendritic, sometimes mixed with simple ones);

petals oblanceolate, 0.7–1 × 0.1–0.2 mm;

median filaments 1.5–2 mm;

anthers 0.2–0.3 mm.

Fruiting pedicels

divaricate to ascending, (often recurved in age), slender, (3–)4–9(–13) mm.

divaricate-ascending, straight, 4–7(–11) mm.

Fruits

erect to widely spreading, narrowly linear, slightly torulose, (9–)14–30(–34) × 0.6–1.1 mm, (usually terete, rarely slightly flattened, often curved inward);

valves each with obscure midvein;

septum not veined;

ovules 30–62 per ovary;

style obsolete, 0.07–0.3 mm, glabrous.

divaricate to erect, linear to oblong, not torulose, 3–8(–10) × 1–1.2 mm;

valves each with distinct midvein;

septum not veined;

ovules 10–28 per ovary;

style obsolete to 0.1 mm, glabrous.

Seeds

uniseriate, light brown, narrowly oblong, 1–1.5 × 0.3–0.5 mm.

biseriate, light brown, ellipsoid, 0.5–0.7 × 0.3–0.4 mm.

2n

= 14.

= 42.

Descurainia sophioides

Descurainia brevisiliqua

Phenology Flowering Jun–Sep. Flowering (late Jun-)Jul–Aug(-early Oct).
Habitat Open meadows, eroded peat, roadsides, disturbed and waste sites, rocky outcrops, mining dumps, gravelly grounds, stream banks, gullies Pine and juniper communities, rocky washes, roadsides, grasslands, gravelly mesa
Elevation 0-1000 m (0-3300 ft) 1900-2500 m (6200-8200 ft)
Distribution
from FNA
AK; BC; MB; NT; NU; YT; Asia (Russian Far East, Siberia)
[BONAP county map]
from FNA
AZ; NM; TX
[BONAP county map]
Discussion

R. C. Rollins (1993) reduced Descurainia brevisiliqua to synonymy of the diploid D. obtusa. This hexaploid species differs from the latter in having biseriate (versus uniseriate) seeds and glabrous (versus densely to moderately pubescent) fruits. Molecular data (B. E. Goodson 2007) show that it is consistently separate from, and unrelated to, D. obtusa, instead being affiliated with D. incana and D. incisa. Descurainia brevisiliqua differs from the latter two species by having biseriate (versus uniseriate) seeds and oblong, non-torulose fruits that, unlike those of D. incana, are not appressed to rachises. Although the fruits of D. brevisiliqua resemble those of some D. pinnata subspecies, it can be readily distinguished from the latter by its tall, strict growth habit with numerous short branches in the distal part of the plant, more linear leaf segments, stems that are usually purple, and late–summer flowering period.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 529. FNA vol. 7, p. 521.
Parent taxa Brassicaceae > tribe Descurainieae > Descurainia Brassicaceae > tribe Descurainieae > Descurainia
Sibling taxa
D. adenophora, D. brevisiliqua, D. californica, D. incana, D. incisa, D. kenheilii, D. longepedicellata, D. nelsonii, D. obtusa, D. paradisa, D. pinnata, D. sophia, D. torulosa
D. adenophora, D. californica, D. incana, D. incisa, D. kenheilii, D. longepedicellata, D. nelsonii, D. obtusa, D. paradisa, D. pinnata, D. sophia, D. sophioides, D. torulosa
Synonyms Sisymbrium sophioides, Hesperis arctica, Sisymbrium arcticum, Sophia sophioides D. obtusa subsp. brevisiliqua
Name authority (Fischer ex Hooker) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 86[IV,105]: 316. (1924) (Detling) Al-Shehbaz & Goodson: Harvard Pap. Bot. 12: 421. (2007)
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