Descurainia sophia |
Descurainia torulosa |
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fine-leaf tansy-mustard, fixweed, flixweed, flixweed tansymustard, herb sophia, not much flower, tansy mustard |
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Habit | Annuals; eglandular; sparsely to densely pubescent, sometimes glabrous distally, trichomes dendritic. | Perennials; (short-lived); eglandular; moderately to densely pubescent throughout, trichomes dendritic. |
Stems | erect, unbranched or branched distally, (1–)2–7(–10) dm. |
(several to numerous from base), decumbent, usually unbranched, rarely branched distally, 0.4–1.5 dm. |
Basal leaves | petiole 0.1–2(–3) cm; blade 2- or 3-pinnate, ovate or oblong to obovate in outline, to 15 cm, lateral lobes linear or oblong, (to 10 × 2 mm), margins entire. |
petiole 0.3–1.4 cm; blade pinnate, oblanceolate in outline, 0.9–3.5 cm, lateral lobes (3–5 pairs), oblanceolate to oblong, (2–5 × 0.5–1.5 mm), margins entire. |
Cauline leaves | sessile or shortly petiolate; blade smaller distally, distal lobes often narrower, surfaces often glabrous. |
sessile or shortly petiolate; blade smaller distally, distal lobes often narrower, surfaces moderately pubescent. |
Racemes | considerably elongated in fruit. |
considerably elongated in fruit. |
Flowers | sepals erect to ascending, yellowish, oblong, 1.8–2.8 mm, glabrate to sparsely pubescent; petals narrowly oblanceolate, 2–3 × 0.4–0.6 mm; median filaments 2–3 mm; anthers 0.3–0.4 mm. |
sepals ascending, yellowish, oblong, 0.9–1.2 mm, pubescent; petals narrowly oblanceolate, 1.4–2 × 0.4–0.6 mm; median filaments 1.6–2 mm; anthers 0.2–0.3 mm. |
Fruiting pedicels | divaricate to ascending, straight, (5–)8–15(–20) mm. |
erect to erect-ascending, straight, 1.5–3 mm. |
Fruits | divaricate-ascending to erect, narrowly linear, torulose, (12–)15–27(–30) × 0.5–0.8(–1) mm, (straight or curved upward); valves each with distinct midvein; septum with a broad central longitudinal band appearing as 2 or 3 veins; ovules 20–48 per ovary; style obsolete, 0.05–0.2 mm, glabrous. |
erect, (subappressed to rachis basally), narrowly linear, strongly torulose, 6–15 × 0.6–0.8 mm, (curved outward distally); valves each with obscure midvein, (pubescent); septum not veined; ovules 10–18 per ovary; style obsolete, 0.1–0.3 mm, sparsely pubescent. |
Seeds | uniseriate, reddish brown, oblong, 0.7–1.3 × 0.3–0.6 mm. |
uniseriate, reddish brown, oblong, 1–1.3 × 0.5–0.6 mm. |
2n | = 28. |
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Descurainia sophia |
Descurainia torulosa |
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Phenology | Flowering Mar–Jul. | Flowering Jun–Jul. |
Habitat | Roadsides, waste places, disturbed sites, railroad embankments, hillsides, mountain slopes, canyon bottoms, stream banks, fields, lawns, pastures, deserts, sagebrush and pinyon-juniper communities | Rocky slopes at bases of cliffs |
Elevation | 0-3000 m [0-9800 ft] | ca. 3100 m [ca. 10200 ft] |
Distribution |
AK; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; ME; MI; MN; MO; MT; NC; ND; NE; NH; NM; NV; NY; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, South Africa, Australia]
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WY |
Discussion | Deviant chromosome counts (e.g., 2n = 12, 14, 20, 38; see R. C. Rollins 1993, N. H. Holmgren 2005b, S. I. Warwick and I. A. Al-Shehbaz 2006) are most certainly erroneous, and the species appears to be exclusively tetraploid based on x = 7. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Descurainia torulosa is known from Fremont, Park, Sweetwater, and Teton counties. It was studied by J. S. Bricker et al. (2000), who concluded that its status and relationship to D. incana remain unresolved as they were unable to find a set of morphological characters that consistently separated the two taxa. We believe that D. torulosa should be maintained, and that it is easily distinguished from D. incana by having fruits strongly torulose (versus weakly or not torulose), pubescent (versus glabrous), and distinctly curved (versus straight) distally, sparsely pubescent (versus glabrous) styles, fruit valves each with obscure (versus distinct) midvein, and stems decumbent, unbranched, and often several (versus stems erect, branched, and simple) from the base. Descurainia torulosa is in the Center for Plant Conservation’s National Database of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 528. | FNA vol. 7, p. 529. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Sisymbrium sophia, Hesperis sophia, Sisymbrium parviflorum, Sophia parviflora | |
Name authority | (Linnaeus) Webb ex Prantl: in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 55(III,2): 192. (1891) | Rollins: J. Arnold Arbor. 64: 499. (1983) |
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