FNA: Delphinium umbraculorum vs. Delphinium polycladon

	Delphinium umbraculorum	Delphinium polycladon
	umbrella larkspur	high mountain larkspur, mountain marsh larkspur
Stems	40-70(-90) cm; base often reddish, glabrous or puberulent.	(20-)60-100(-160) cm; base reddish or not, glabrous.
Leaves	blade round to pentagonal, 1.5-4 × 2-6 cm, nearly glabrous; ultimate lobes 3-13, width 3-20 mm (basal), 1-8 mm (cauline).	blade round to pentagonal, 1.5-7 × 2-14 cm, glabrous; ultimate lobes 3-12, width 4-30(-45) mm (basal), 3-30 mm (cauline).
Inflorescences	(5-)10-25(-45)-flowered, open, narrowly pyramidal; pedicel 0.5-3(-7) cm, glabrous to puberulent; bracteoles 3-7 mm from flowers, green, linear, 3-6 mm, puberulent.	3-15(-35)-flowered, open, often ± secund; pedicel 1-4(-15) cm, glabrous to puberulent; bracteoles 2-8(-37) mm from flowers, green, linear, 4-7(-11) mm, nearly glabrous.
Flowers	sepals dark blue, puberulent, lateral sepals spreading, 9-16 × 4-7 mm, spurs gently upcurved, ascending 30-45° above horizontal, 8-14 mm; lower petal blades elevated, exposing stamens, 3.5-6 mm, clefts 0.5-1.5 mm; hairs densest near junction of blade and claw above base of cleft, centered or on inner lobes, white.	sepals bluish purple, nearly glabrous, lateral sepals spreading, (10-)12-18 × 7-10 mm, spurs usually downcurved, ca. 30° below horizontal, 11-22 mm; lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 1-2 mm; hairs mostly near base of cleft on inner lobes, yellow, sometimes white.
Fruits	9-16(-19) mm, 2.5-3(-4) times longer than wide, puberulent.	13-20 mm, 3.5-4 times longer than wide, puberulent.
Seeds	seed coat cells brick-shaped, cell margins straight, surfaces smooth.	unwinged; seed coat cells with surfaces roughened.
2n	= 16.	= 16.

	Delphinium umbraculorum	Delphinium polycladon
Phenology	Flowering late spring–early summer.	Flowering summer–early autumn.
Habitat	Slopes in oak forests	Wet sites near springs, streamsides, bogs, and wet talus
Elevation	400-1600 m (1300-5200 ft)	2200-3600 m (7200-11800 ft)
Distribution	CA [WildflowerSearch map] [BONAP county map]	CA; NV [WildflowerSearch map] [BONAP county map]
Discussion	Delphinium umbraculorum is most often confused with D. patens subsp. hepaticoideum; refer to discussion of that taxon for distinguishing features. Hybrids occur with D. parryi and D. patens subsp. montanum. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium polycladon hybridizes with D. depauperatum and D. glaucum. Plants of D. polycladon are extremely variable. Individuals from very rocky, thin-soiled, sunny sites at higher elevations tend to be quite compact; they show the features of the species in a dwarfed state. Proximal internodes are especially shortened. Plants from areas of deeper soil (high or low elevations), especially those growing among shrubs, usually are much taller, with elongate proximal internodes, and other vegetative parts proportionally larger. Shorter plants may be confused with D. depauperatum or D. nuttallianum; see discussion under those species for distinguishing features. Taller plants may be confused with D. glaucum; they can be distinguished by their leaves predominately on proximal part of stem, sigmoid pedicel, and fewer flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Subscaposa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. scopulorum var. luporum
Name authority	H. F. Lewis & Epling: Brittonia 8: 19. (1954)	Eastwood: Bull. Torrey Bot. Club 28: 669. (1901)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium polycladon

umbrella larkspur

high mountain larkspur, mountain marsh larkspur

Stems

40-70(-90) cm;

base often reddish, glabrous or puberulent.

(20-)60-100(-160) cm;

base reddish or not, glabrous.

Leaves

blade round to pentagonal, 1.5-4 × 2-6 cm, nearly glabrous; ultimate lobes 3-13, width 3-20 mm (basal), 1-8 mm (cauline).

blade round to pentagonal, 1.5-7 × 2-14 cm, glabrous; ultimate lobes 3-12, width 4-30(-45) mm (basal), 3-30 mm (cauline).

Inflorescences

(5-)10-25(-45)-flowered, open, narrowly pyramidal;

pedicel 0.5-3(-7) cm, glabrous to puberulent;

bracteoles 3-7 mm from flowers, green, linear, 3-6 mm, puberulent.

3-15(-35)-flowered, open, often ± secund;

pedicel 1-4(-15) cm, glabrous to puberulent;

bracteoles 2-8(-37) mm from flowers, green, linear, 4-7(-11) mm, nearly glabrous.

Flowers

sepals dark blue, puberulent, lateral sepals spreading, 9-16 × 4-7 mm, spurs gently upcurved, ascending 30-45° above horizontal, 8-14 mm;

lower petal blades elevated, exposing stamens, 3.5-6 mm, clefts 0.5-1.5 mm;

hairs densest near junction of blade and claw above base of cleft, centered or on inner lobes, white.

sepals bluish purple, nearly glabrous, lateral sepals spreading, (10-)12-18 × 7-10 mm, spurs usually downcurved, ca. 30° below horizontal, 11-22 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 1-2 mm;

hairs mostly near base of cleft on inner lobes, yellow, sometimes white.

Fruits

9-16(-19) mm, 2.5-3(-4) times longer than wide, puberulent.

13-20 mm, 3.5-4 times longer than wide, puberulent.

Seeds

seed coat cells brick-shaped, cell margins straight, surfaces smooth.

unwinged;

seed coat cells with surfaces roughened.

= 16.

Delphinium polycladon

Phenology

Flowering late spring–early summer.

Flowering summer–early autumn.

Habitat

Slopes in oak forests

Wet sites near springs, streamsides, bogs, and wet talus

Elevation

400-1600 m (1300-5200 ft)

2200-3600 m (7200-11800 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

CA; NV

[WildflowerSearch map]

[BONAP county map]

Discussion

Delphinium umbraculorum is most often confused with D. patens subsp. hepaticoideum; refer to discussion of that taxon for distinguishing features. Hybrids occur with D. parryi and D. patens subsp. montanum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium polycladon hybridizes with D. depauperatum and D. glaucum. Plants of D. polycladon are extremely variable. Individuals from very rocky, thin-soiled, sunny sites at higher elevations tend to be quite compact; they show the features of the species in a dwarfed state. Proximal internodes are especially shortened. Plants from areas of deeper soil (high or low elevations), especially those growing among shrubs, usually are much taller, with elongate proximal internodes, and other vegetative parts proportionally larger. Shorter plants may be confused with D. depauperatum or D. nuttallianum; see discussion under those species for distinguishing features. Taller plants may be confused with D. glaucum; they can be distinguished by their leaves predominately on proximal part of stem, sigmoid pedicel, and fewer flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Subscaposa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. scopulorum var. luporum

Name authority

H. F. Lewis & Epling: Brittonia 8: 19. (1954)

Eastwood: Bull. Torrey Bot. Club 28: 669. (1901)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Delphinium umbraculorum

Delphinium polycladon

Delphinium umbraculorum

Delphinium polycladon