FNA: Delphinium uliginosum vs. Delphinium polycladon

	Delphinium uliginosum	Delphinium polycladon
	swamp larkspur	high mountain larkspur, mountain marsh larkspur
Stems	10-30(-70) cm; base reddish or not, nearly glabrous.	(20-)60-100(-160) cm; base reddish or not, glabrous.
Leaves	blade obdeltoid, apically several parted, 1-8 × 1-7 cm, ± fleshy, glabrous; ultimate lobes 0-3, width 3-20 mm (cauline only); margins of basal leaf, measured less than 1 cm from blade base, demarcating less than 90° of arc when leaf laid flat.	blade round to pentagonal, 1.5-7 × 2-14 cm, glabrous; ultimate lobes 3-12, width 4-30(-45) mm (basal), 3-30 mm (cauline).
Inflorescences	5-20(-48)-flowered, ± open; pedicel 0.3-3(-10) cm, glabrous to puberulent; bracteoles 2-3(-5) mm from flowers, green to blue, lanceolate-linear, 3-4(-7) mm, puberulent.	3-15(-35)-flowered, open, often ± secund; pedicel 1-4(-15) cm, glabrous to puberulent; bracteoles 2-8(-37) mm from flowers, green, linear, 4-7(-11) mm, nearly glabrous.
Flowers	sepals dark blue, nearly glabrous, lateral sepals spreading, 9-15 × 5-8 mm, spurs usually upcurved, ascending 30-45° above horizontal, 10-14 mm; lower petal blades slightly elevated, ± exposing stamens, 4-5 mm, clefts 2-3 mm; hairs centered, densest on inner lobe above base of cleft, also on margins, white.	sepals bluish purple, nearly glabrous, lateral sepals spreading, (10-)12-18 × 7-10 mm, spurs usually downcurved, ca. 30° below horizontal, 11-22 mm; lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 1-2 mm; hairs mostly near base of cleft on inner lobes, yellow, sometimes white.
Fruits	10-18 mm, 4.1-4.5 times longer than wide, puberulent.	13-20 mm, 3.5-4 times longer than wide, puberulent.
Seeds	seed coat cells with surfaces bumpy or wavy.	unwinged; seed coat cells with surfaces roughened.
2n	= 16.	= 16.

	Delphinium uliginosum	Delphinium polycladon
Phenology	Flowering late spring–early summer.	Flowering summer–early autumn.
Habitat	Serpentine streamsides, chaparral, grassland	Wet sites near springs, streamsides, bogs, and wet talus
Elevation	400-600 m (1300-2000 ft)	2200-3600 m (7200-11800 ft)
Distribution	CA [WildflowerSearch map] [BONAP county map]	CA; NV [WildflowerSearch map] [BONAP county map]
Discussion	Although some populations are large, Delphinium uliginosum is very local. Hybrids with D. hesperium subsp. pallescens have been seen. Delphinium uliginosum is a very distinctive species, not likely to be confused with any other. The fan-shaped, slightly dissected leaves are apparently unique in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium polycladon hybridizes with D. depauperatum and D. glaucum. Plants of D. polycladon are extremely variable. Individuals from very rocky, thin-soiled, sunny sites at higher elevations tend to be quite compact; they show the features of the species in a dwarfed state. Proximal internodes are especially shortened. Plants from areas of deeper soil (high or low elevations), especially those growing among shrubs, usually are much taller, with elongate proximal internodes, and other vegetative parts proportionally larger. Shorter plants may be confused with D. depauperatum or D. nuttallianum; see discussion under those species for distinguishing features. Taller plants may be confused with D. glaucum; they can be distinguished by their leaves predominately on proximal part of stem, sigmoid pedicel, and fewer flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Depauperata	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. scopulorum var. luporum
Name authority	Curran: Proc. Calif. Acad. Sci. 1: 151. (1885)	Eastwood: Bull. Torrey Bot. Club 28: 669. (1901)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium polycladon

swamp larkspur

high mountain larkspur, mountain marsh larkspur

Stems

10-30(-70) cm;

base reddish or not, nearly glabrous.

(20-)60-100(-160) cm;

base reddish or not, glabrous.

Leaves

blade obdeltoid, apically several parted, 1-8 × 1-7 cm, ± fleshy, glabrous; ultimate lobes 0-3, width 3-20 mm (cauline only);

margins of basal leaf, measured less than 1 cm from blade base, demarcating less than 90° of arc when leaf laid flat.

blade round to pentagonal, 1.5-7 × 2-14 cm, glabrous; ultimate lobes 3-12, width 4-30(-45) mm (basal), 3-30 mm (cauline).

Inflorescences

5-20(-48)-flowered, ± open;

pedicel 0.3-3(-10) cm, glabrous to puberulent;

bracteoles 2-3(-5) mm from flowers, green to blue, lanceolate-linear, 3-4(-7) mm, puberulent.

3-15(-35)-flowered, open, often ± secund;

pedicel 1-4(-15) cm, glabrous to puberulent;

bracteoles 2-8(-37) mm from flowers, green, linear, 4-7(-11) mm, nearly glabrous.

Flowers

sepals dark blue, nearly glabrous, lateral sepals spreading, 9-15 × 5-8 mm, spurs usually upcurved, ascending 30-45° above horizontal, 10-14 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-5 mm, clefts 2-3 mm;

hairs centered, densest on inner lobe above base of cleft, also on margins, white.

sepals bluish purple, nearly glabrous, lateral sepals spreading, (10-)12-18 × 7-10 mm, spurs usually downcurved, ca. 30° below horizontal, 11-22 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 1-2 mm;

hairs mostly near base of cleft on inner lobes, yellow, sometimes white.

Fruits

10-18 mm, 4.1-4.5 times longer than wide, puberulent.

13-20 mm, 3.5-4 times longer than wide, puberulent.

Seeds

seed coat cells with surfaces bumpy or wavy.

unwinged;

seed coat cells with surfaces roughened.

= 16.

Delphinium polycladon

Phenology

Flowering late spring–early summer.

Flowering summer–early autumn.

Habitat

Serpentine streamsides, chaparral, grassland

Wet sites near springs, streamsides, bogs, and wet talus

Elevation

400-600 m (1300-2000 ft)

2200-3600 m (7200-11800 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

CA; NV

[WildflowerSearch map]

[BONAP county map]

Discussion

Although some populations are large, Delphinium uliginosum is very local. Hybrids with D. hesperium subsp. pallescens have been seen.

Delphinium uliginosum is a very distinctive species, not likely to be confused with any other. The fan-shaped, slightly dissected leaves are apparently unique in the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium polycladon hybridizes with D. depauperatum and D. glaucum. Plants of D. polycladon are extremely variable. Individuals from very rocky, thin-soiled, sunny sites at higher elevations tend to be quite compact; they show the features of the species in a dwarfed state. Proximal internodes are especially shortened. Plants from areas of deeper soil (high or low elevations), especially those growing among shrubs, usually are much taller, with elongate proximal internodes, and other vegetative parts proportionally larger. Shorter plants may be confused with D. depauperatum or D. nuttallianum; see discussion under those species for distinguishing features. Taller plants may be confused with D. glaucum; they can be distinguished by their leaves predominately on proximal part of stem, sigmoid pedicel, and fewer flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Depauperata

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. scopulorum var. luporum

Name authority

Curran: Proc. Calif. Acad. Sci. 1: 151. (1885)

Eastwood: Bull. Torrey Bot. Club 28: 669. (1901)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Delphinium uliginosum

Delphinium polycladon

Delphinium uliginosum

Delphinium polycladon