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swamp larkspur

Greene's larkspur, meadow larkspur, pine forest larkspur, slender or Greene's larkspur

Stems

10-30(-70) cm;

base reddish or not, nearly glabrous.

(15-)30-50(-80) cm;

base reddish, nearly glabrous.

Leaves

blade obdeltoid, apically several parted, 1-8 × 1-7 cm, ± fleshy, glabrous; ultimate lobes 0-3, width 3-20 mm (cauline only);

margins of basal leaf, measured less than 1 cm from blade base, demarcating less than 90° of arc when leaf laid flat.

blade round to pentagonal, 1.5-4 × 3-7 cm, nearly glabrous; ultimate lobes 3-7, distinctly wedge-shaped, usually 5 or fewer extending 3/5 distance to petiole, width 5-20 mm (basal), 1-15 mm (cauline), widest in distal 1/2.

Inflorescences

5-20(-48)-flowered, ± open;

pedicel 0.3-3(-10) cm, glabrous to puberulent;

bracteoles 2-3(-5) mm from flowers, green to blue, lanceolate-linear, 3-4(-7) mm, puberulent.

5-20(-38)-flowered;

pedicel spreading from rachis at nearly 90°, 1-3(-4) cm, glabrous or glandular-pubescent;

bracteoles (7-)11-19 mm from flowers, blue or green, linear, 2-5 mm, puberulent to glabrous.

Flowers

sepals dark blue, nearly glabrous, lateral sepals spreading, 9-15 × 5-8 mm, spurs usually upcurved, ascending 30-45° above horizontal, 10-14 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-5 mm, clefts 2-3 mm;

hairs centered, densest on inner lobe above base of cleft, also on margins, white.

sepals dark bluish purple to pink or white, usually retaining color upon drying, glabrous, lateral sepals reflexed, 6-10(-13) × 3-6 mm, spurs often curved upward, within 30° above or below horizontal, 8-12(-14) mm;

lower petal blades elevated, exposing stamens, 3-5 mm, clefts 1-3 mm;

hairs almost exclusively near base of cleft, centered or mostly on inner lobes, usually yellow.

Fruits

10-18 mm, 4.1-4.5 times longer than wide, puberulent.

8-16 mm, 3-3.5 times longer than wide, glabrous to glandular-puberulent.

Seeds

seed coat cells with surfaces bumpy or wavy.

unwinged;

seed coats ± pitted, cell surfaces roughened.

2n

= 16.

Delphinium uliginosum

Delphinium gracilentum

Phenology Flowering late spring–early summer. Flowering spring–early summer.
Habitat Serpentine streamsides, chaparral, grassland Open coniferous forest
Elevation 400-600 m (1300-2000 ft) 150-2700 m (500-8900 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Although some populations are large, Delphinium uliginosum is very local. Hybrids with D. hesperium subsp. pallescens have been seen.

Delphinium uliginosum is a very distinctive species, not likely to be confused with any other. The fan-shaped, slightly dissected leaves are apparently unique in the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium gracilentum hybridizes with D. patens subsp. patens in the northern Sierra Nevada foothills and is very similar to that species, making hybrids difficult to discern. While D. gracilentum and D. patens are easily distinguished in most of their ranges, morphologic distinctions between the two taxa are blurred in the northern Sierra Nevada foothills region, particularly in Butte County, California. Coniferous woods are preferred by D. gracilentum; D. patens subspp. patens and hepaticoideum are more often found in broadleaf woods. The former species has more widely spreading pedicels than the latter, and D. gracilentum usually has wider leaf lobes than D. patens subsp. patens. In the southern Sierra Nevada, D. gracilentum may come in contact with D. patens subsp. montanum. Though hybrids are not common, some gene flow has apparently occurred.

Sepal color phases are not stable and considerable variation occurs within populations. The type specimen of Delphinium gracilentum represents the northern, lower elevation, nonglandular, dark-flowered phase. The type specimen of D. gracilentum forma versicolor Ewan differs only by its pink or white flowers. A limited range of intermediate colors occurs, and populations may be made up of plants of a single color or several different colors. The type specimen of D. greenei Eastwood represents the southern, higher elevation, glandular (at least on pedicels) expression. The type specimen includes representatives of dark- and light-flowered individuals of this phase. The type specimen of D. gracilentum forma versicolor (not seen by the author) is the "albino" phase referred to by Greene in his description of D. gracilentum. Several of the paratypes cited by Ewan have been seen, as have a number of individuals in natural populations.

Delphinium gracilentum has been confused with D. patens or D. nuttallianum. Delphinium gracilentum may be distinguished from D. nuttallianum by its wider leaf lobes, smaller fruits, and more elongate inflorescences, and from D. patens by its wider leaf lobes, more open inflorescences, and usually shorter fruits.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Depauperata Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms D. patens subsp. greenei, D. pratense
Name authority Curran: Proc. Calif. Acad. Sci. 1: 151. (1885) Greene: Pittonia 3: 15. (1896)
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