FNA: Delphinium trolliifolium vs. Delphinium polycladon

	Delphinium trolliifolium	Delphinium polycladon
	Columbian larkspur, cow-poison, poison delphinium, poison larkspur	high mountain larkspur, mountain marsh larkspur
Stems	(40-)60-120(-180) cm; base usually reddish, glabrous to puberulent.	(20-)60-100(-160) cm; base reddish or not, glabrous.
Leaves	blade ± pentagonal, 4-8 × 7-16 cm, margins ± incised, nearly glabrous; ultimate lobes 0-9, width 15-30 mm (basal), 5-20 mm (cauline), widest at middle or in proximal 1/2.	blade round to pentagonal, 1.5-7 × 2-14 cm, glabrous; ultimate lobes 3-12, width 4-30(-45) mm (basal), 3-30 mm (cauline).
Inflorescences	(5-)14-40(-75)-flowered, ± open, at least 2 times longer than wide; pedicel 1-4(-9) cm, puberulent to glabrous; bracteoles (2-)6-12 mm from flowers, green, linear, 5-9(-14) mm, puberulent.	3-15(-35)-flowered, open, often ± secund; pedicel 1-4(-15) cm, glabrous to puberulent; bracteoles 2-8(-37) mm from flowers, green, linear, 4-7(-11) mm, nearly glabrous.
Flowers	sepals dark blue, glabrous, lateral sepals spreading, (8-)14-21 × 5-9 mm, spurs straight or downcurved at apex, within 20° of horizontal, (10-)16-23 mm; lower petal blades covering stamens, 5-10 mm, clefts 1.5-3 mm; hairs sparse, mostly near junction of blade and claw, centered or on inner lobes, well dispersed, yellow.	sepals bluish purple, nearly glabrous, lateral sepals spreading, (10-)12-18 × 7-10 mm, spurs usually downcurved, ca. 30° below horizontal, 11-22 mm; lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 1-2 mm; hairs mostly near base of cleft on inner lobes, yellow, sometimes white.
Fruits	(15-)23-34 mm, 3.8-5.5 times longer than wide, glabrous.	13-20 mm, 3.5-4 times longer than wide, puberulent.
Seeds	unwinged; seed coats smooth.	unwinged; seed coat cells with surfaces roughened.
2n	= 16.	= 16.

	Delphinium trolliifolium	Delphinium polycladon
Phenology	Flowering spring.	Flowering summer–early autumn.
Habitat	Oak woods, coastal chaparral, wet woodlands	Wet sites near springs, streamsides, bogs, and wet talus
Elevation	30-1100 m (100-3600 ft)	2200-3600 m (7200-11800 ft)
Distribution	CA; OR [WildflowerSearch map] [BONAP county map]	CA; NV [WildflowerSearch map] [BONAP county map]
Discussion	Delphinium trolliifolium occurs in the northern Coast Range of California, the Columbia River Valley to just east of Mt. Hood, and the Willamette Valley of Oregon upstream to Lane County. California plants differ somewhat from Oregon plants in pubescence patterns and habitat preferences. Further study may show that two entities are involved here. Hybrids between Delphinium trolliifolium and D. decorum, D. menziesii subsp. pallidum (D. ×pavonaceum Ewan, Peacock larkspur), D. nudicaule, D. nuttallianum, and D. nuttallii are known. Delphinium trolliifolium is likely to be confused only with D. bakeri. Refer to discussion under that species for differences. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium polycladon hybridizes with D. depauperatum and D. glaucum. Plants of D. polycladon are extremely variable. Individuals from very rocky, thin-soiled, sunny sites at higher elevations tend to be quite compact; they show the features of the species in a dwarfed state. Proximal internodes are especially shortened. Plants from areas of deeper soil (high or low elevations), especially those growing among shrubs, usually are much taller, with elongate proximal internodes, and other vegetative parts proportionally larger. Shorter plants may be confused with D. depauperatum or D. nuttallianum; see discussion under those species for distinguishing features. Taller plants may be confused with D. glaucum; they can be distinguished by their leaves predominately on proximal part of stem, sigmoid pedicel, and fewer flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3.	FNA vol. 3.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. scopulorum var. luporum
Name authority	A. Gray: Proc. Amer. Acad. Arts 8: 375. (1872)	Eastwood: Bull. Torrey Bot. Club 28: 669. (1901)
Web links	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium polycladon

Columbian larkspur, cow-poison, poison delphinium, poison larkspur

high mountain larkspur, mountain marsh larkspur

Stems

(40-)60-120(-180) cm;

base usually reddish, glabrous to puberulent.

(20-)60-100(-160) cm;

base reddish or not, glabrous.

Leaves

blade ± pentagonal, 4-8 × 7-16 cm, margins ± incised, nearly glabrous; ultimate lobes 0-9, width 15-30 mm (basal), 5-20 mm (cauline), widest at middle or in proximal 1/2.

blade round to pentagonal, 1.5-7 × 2-14 cm, glabrous; ultimate lobes 3-12, width 4-30(-45) mm (basal), 3-30 mm (cauline).

Inflorescences

(5-)14-40(-75)-flowered, ± open, at least 2 times longer than wide;

pedicel 1-4(-9) cm, puberulent to glabrous;

bracteoles (2-)6-12 mm from flowers, green, linear, 5-9(-14) mm, puberulent.

3-15(-35)-flowered, open, often ± secund;

pedicel 1-4(-15) cm, glabrous to puberulent;

bracteoles 2-8(-37) mm from flowers, green, linear, 4-7(-11) mm, nearly glabrous.

Flowers

sepals dark blue, glabrous, lateral sepals spreading, (8-)14-21 × 5-9 mm, spurs straight or downcurved at apex, within 20° of horizontal, (10-)16-23 mm;

lower petal blades covering stamens, 5-10 mm, clefts 1.5-3 mm;

hairs sparse, mostly near junction of blade and claw, centered or on inner lobes, well dispersed, yellow.

sepals bluish purple, nearly glabrous, lateral sepals spreading, (10-)12-18 × 7-10 mm, spurs usually downcurved, ca. 30° below horizontal, 11-22 mm;

lower petal blades slightly elevated, ± exposing stamens, 4-6 mm, clefts 1-2 mm;

hairs mostly near base of cleft on inner lobes, yellow, sometimes white.

Fruits

(15-)23-34 mm, 3.8-5.5 times longer than wide, glabrous.

13-20 mm, 3.5-4 times longer than wide, puberulent.

Seeds

unwinged;

seed coats smooth.

unwinged;

seed coat cells with surfaces roughened.

= 16.

Delphinium polycladon

Phenology

Flowering spring.

Flowering summer–early autumn.

Habitat

Oak woods, coastal chaparral, wet woodlands

Wet sites near springs, streamsides, bogs, and wet talus

Elevation

30-1100 m (100-3600 ft)

2200-3600 m (7200-11800 ft)

Distribution

CA; OR

[WildflowerSearch map]

[BONAP county map]

CA; NV

[WildflowerSearch map]

[BONAP county map]

Discussion

Delphinium trolliifolium occurs in the northern Coast Range of California, the Columbia River Valley to just east of Mt. Hood, and the Willamette Valley of Oregon upstream to Lane County. California plants differ somewhat from Oregon plants in pubescence patterns and habitat preferences. Further study may show that two entities are involved here.

Hybrids between Delphinium trolliifolium and D. decorum, D. menziesii subsp. pallidum (D. ×pavonaceum Ewan, Peacock larkspur), D. nudicaule, D. nuttallianum, and D. nuttallii are known. Delphinium trolliifolium is likely to be confused only with D. bakeri. Refer to discussion under that species for differences.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium polycladon hybridizes with D. depauperatum and D. glaucum. Plants of D. polycladon are extremely variable. Individuals from very rocky, thin-soiled, sunny sites at higher elevations tend to be quite compact; they show the features of the species in a dwarfed state. Proximal internodes are especially shortened. Plants from areas of deeper soil (high or low elevations), especially those growing among shrubs, usually are much taller, with elongate proximal internodes, and other vegetative parts proportionally larger. Shorter plants may be confused with D. depauperatum or D. nuttallianum; see discussion under those species for distinguishing features. Taller plants may be confused with D. glaucum; they can be distinguished by their leaves predominately on proximal part of stem, sigmoid pedicel, and fewer flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. scopulorum var. luporum

Name authority

A. Gray: Proc. Amer. Acad. Arts 8: 375. (1872)

Eastwood: Bull. Torrey Bot. Club 28: 669. (1901)

Web links

Local floras: CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog.
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Delphinium trolliifolium

Delphinium polycladon

Delphinium trolliifolium

Delphinium polycladon