FNA: Delphinium trolliifolium vs. Delphinium inopinum

	Delphinium trolliifolium	Delphinium inopinum
	Columbian larkspur, cow-poison, poison delphinium, poison larkspur	unexpected larkspur
Stems	(40-)60-120(-180) cm; base usually reddish, glabrous to puberulent.	70-110(-150) cm; base reddish or not, glabrous, often glaucous.
Leaves	blade ± pentagonal, 4-8 × 7-16 cm, margins ± incised, nearly glabrous; ultimate lobes 0-9, width 15-30 mm (basal), 5-20 mm (cauline), widest at middle or in proximal 1/2.	blade ± pentagonal, 1-5 × 1.5-7 cm, glabrous; ultimate lobes 3-9, width 5-28 mm (basal), 3-18 mm (cauline).
Inflorescences	(5-)14-40(-75)-flowered, ± open, at least 2 times longer than wide; pedicel 1-4(-9) cm, puberulent to glabrous; bracteoles (2-)6-12 mm from flowers, green, linear, 5-9(-14) mm, puberulent.	20-35(-51)-flowered, usually dense; pedicel 0.3-1.5(-2.5) cm, glabrous; bracteoles 2-4 mm from flowers, green, linear, 1-2(-4) mm, nearly glabrous.
Flowers	sepals dark blue, glabrous, lateral sepals spreading, (8-)14-21 × 5-9 mm, spurs straight or downcurved at apex, within 20° of horizontal, (10-)16-23 mm; lower petal blades covering stamens, 5-10 mm, clefts 1.5-3 mm; hairs sparse, mostly near junction of blade and claw, centered or on inner lobes, well dispersed, yellow.	sepals white to light blue, glabrous, lateral sepals spreading to forward pointing, 8-12 × 3-5 mm, spurs straight to gently upcurved, ascending 30-60° above horizontal, 9-12 mm; lower petal blades slightly elevated, ± exposing stamens, 3-5 mm, clefts 1-2 mm; hairs centered, densest near base of cleft, white.
Fruits	(15-)23-34 mm, 3.8-5.5 times longer than wide, glabrous.	12-20 mm, 2.6-4 times longer than wide, glabrous.
Seeds	unwinged; seed coats smooth.	wing-margined; seed coat cells surfaces smooth.
2n	= 16.

	Delphinium trolliifolium	Delphinium inopinum
Phenology	Flowering spring.	Flowering summer.
Habitat	Oak woods, coastal chaparral, wet woodlands	Rock outcrops in open coniferous woods
Elevation	30-1100 m [100-3600 ft]	2200-2800 m [7200-9200 ft]
Distribution	CA; OR [WildflowerSearch map] [BONAP county map]	CA [BONAP county map]
Discussion	Delphinium trolliifolium occurs in the northern Coast Range of California, the Columbia River Valley to just east of Mt. Hood, and the Willamette Valley of Oregon upstream to Lane County. California plants differ somewhat from Oregon plants in pubescence patterns and habitat preferences. Further study may show that two entities are involved here. Hybrids between Delphinium trolliifolium and D. decorum, D. menziesii subsp. pallidum (D. ×pavonaceum Ewan, Peacock larkspur), D. nudicaule, D. nuttallianum, and D. nuttallii are known. Delphinium trolliifolium is likely to be confused only with D. bakeri. Refer to discussion under that species for differences. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Delphinium inopinum is apparently endemic to a white metamorphic rock substrate in the Piute Mountains and southern Sierra Nevada. It is not known to hybridize with any other species, although D. patens subsp. montanum has been collected (when both were flowering) within 1 km of D. inopinum and probably occurs much closer. Delphinium inopinum is often confused with D. parishii subsp. pallidum and superficially resembles some white-flowered individuals of D. hansenii, as well as D. gypsophilum and D. hesperium subsp. pallescens. The massive roots with prominent buds readily distinguish D. inopinum from all of these. In addition, the pubescence found on D. hansenii will separate it from the glabrous D. inopinum. Leaves are rarely seen at anthesis near the base of the stem in D. hesperium subsp. pallescens; they are present in D. inopinum. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 3. Treatment author: Michael J. Warnock.	FNA vol. 3. Treatment author: Michael J. Warnock.
Parent taxa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa	Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex
Sibling taxa	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum	D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Synonyms		D. parishii var. inopinum
Name authority	A. Gray: Proc. Amer. Acad. Arts 8: 375. (1872)	(Jepson) H. F. Lewis & Epling: Brittonia 8: 11. (1954)
Web links	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Delphinium inopinum

Columbian larkspur, cow-poison, poison delphinium, poison larkspur

unexpected larkspur

Stems

(40-)60-120(-180) cm;

base usually reddish, glabrous to puberulent.

70-110(-150) cm;

base reddish or not, glabrous, often glaucous.

Leaves

blade ± pentagonal, 4-8 × 7-16 cm, margins ± incised, nearly glabrous; ultimate lobes 0-9, width 15-30 mm (basal), 5-20 mm (cauline), widest at middle or in proximal 1/2.

blade ± pentagonal, 1-5 × 1.5-7 cm, glabrous; ultimate lobes 3-9, width 5-28 mm (basal), 3-18 mm (cauline).

Inflorescences

(5-)14-40(-75)-flowered, ± open, at least 2 times longer than wide;

pedicel 1-4(-9) cm, puberulent to glabrous;

bracteoles (2-)6-12 mm from flowers, green, linear, 5-9(-14) mm, puberulent.

20-35(-51)-flowered, usually dense;

pedicel 0.3-1.5(-2.5) cm, glabrous;

bracteoles 2-4 mm from flowers, green, linear, 1-2(-4) mm, nearly glabrous.

Flowers

sepals dark blue, glabrous, lateral sepals spreading, (8-)14-21 × 5-9 mm, spurs straight or downcurved at apex, within 20° of horizontal, (10-)16-23 mm;

lower petal blades covering stamens, 5-10 mm, clefts 1.5-3 mm;

hairs sparse, mostly near junction of blade and claw, centered or on inner lobes, well dispersed, yellow.

sepals white to light blue, glabrous, lateral sepals spreading to forward pointing, 8-12 × 3-5 mm, spurs straight to gently upcurved, ascending 30-60° above horizontal, 9-12 mm;

lower petal blades slightly elevated, ± exposing stamens, 3-5 mm, clefts 1-2 mm;

hairs centered, densest near base of cleft, white.

Fruits

(15-)23-34 mm, 3.8-5.5 times longer than wide, glabrous.

12-20 mm, 2.6-4 times longer than wide, glabrous.

Seeds

unwinged;

seed coats smooth.

wing-margined;

seed coat cells surfaces smooth.

= 16.

Delphinium inopinum

Phenology

Flowering spring.

Flowering summer.

Habitat

Oak woods, coastal chaparral, wet woodlands

Rock outcrops in open coniferous woods

Elevation

30-1100 m [100-3600 ft]

2200-2800 m [7200-9200 ft]

Distribution

CA; OR

[WildflowerSearch map]

[BONAP county map]

Discussion

Delphinium trolliifolium occurs in the northern Coast Range of California, the Columbia River Valley to just east of Mt. Hood, and the Willamette Valley of Oregon upstream to Lane County. California plants differ somewhat from Oregon plants in pubescence patterns and habitat preferences. Further study may show that two entities are involved here.

Hybrids between Delphinium trolliifolium and D. decorum, D. menziesii subsp. pallidum (D. ×pavonaceum Ewan, Peacock larkspur), D. nudicaule, D. nuttallianum, and D. nuttallii are known. Delphinium trolliifolium is likely to be confused only with D. bakeri. Refer to discussion under that species for differences.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Delphinium inopinum is apparently endemic to a white metamorphic rock substrate in the Piute Mountains and southern Sierra Nevada. It is not known to hybridize with any other species, although D. patens subsp. montanum has been collected (when both were flowering) within 1 km of D. inopinum and probably occurs much closer. Delphinium inopinum is often confused with D. parishii subsp. pallidum and superficially resembles some white-flowered individuals of D. hansenii, as well as D. gypsophilum and D. hesperium subsp. pallescens. The massive roots with prominent buds readily distinguish D. inopinum from all of these. In addition, the pubescence found on D. hansenii will separate it from the glabrous D. inopinum. Leaves are rarely seen at anthesis near the base of the stem in D. hesperium subsp. pallescens; they are present in D. inopinum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 3. Treatment author: Michael J. Warnock.

Parent taxa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa

Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Multiplex

Sibling taxa

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum

Synonyms

D. parishii var. inopinum

Name authority

A. Gray: Proc. Amer. Acad. Arts 8: 375. (1872)

(Jepson) H. F. Lewis & Epling: Brittonia 8: 11. (1954)

Web links

Local floras: CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog.
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Delphinium trolliifolium

Delphinium inopinum

Delphinium trolliifolium

Delphinium inopinum