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Delphinium trolliifolium

Columbian larkspur, cow-poison, poison delphinium, poison larkspur

Carolina larkspur

Stems

(40-)60-120(-180) cm;

base usually reddish, glabrous to puberulent.

(20-)40-90(-150) cm;

base reddish or not, ± pubescent.

Leaves

blade ± pentagonal, 4-8 × 7-16 cm, margins ± incised, nearly glabrous; ultimate lobes 0-9, width 15-30 mm (basal), 5-20 mm (cauline), widest at middle or in proximal 1/2.

blade round to pentagonal, 1-8 × 2-12 cm, pubescence variable; ultimate lobes 3-29, width 2-10 mm (basal), 1-7 mm (cauline).

Inflorescences

(5-)14-40(-75)-flowered, ± open, at least 2 times longer than wide;

pedicel 1-4(-9) cm, puberulent to glabrous;

bracteoles (2-)6-12 mm from flowers, green, linear, 5-9(-14) mm, puberulent.

(3-)8-27(-94)-flowered;

pedicel (0.4-)0.7-1.8(-5.7) cm, nearly glabrous to glandular;

bracteoles 1-3.5(-6) mm from flowers, green or blue, linear, 2-7 mm, pubescence nearly glabrous to glandular.

Flowers

sepals dark blue, glabrous, lateral sepals spreading, (8-)14-21 × 5-9 mm, spurs straight or downcurved at apex, within 20° of horizontal, (10-)16-23 mm;

lower petal blades covering stamens, 5-10 mm, clefts 1.5-3 mm;

hairs sparse, mostly near junction of blade and claw, centered or on inner lobes, well dispersed, yellow.

sepals purple to blue to white, nearly glabrous, lateral sepals spreading, (7-)9-14(-17) × (3-)3.5-6(-8) mm, spurs ± upcurved, ascending 20-90° above vertical, (9-)11-17(-19) mm;

lower petal blades ± covering stamens, 5-7 mm, cleft 2-4 mm;

hairs centered, densest near base of cleft, white, sometimes blue or yellow.

Fruits

(15-)23-34 mm, 3.8-5.5 times longer than wide, glabrous.

(10-)12.5-18.5(-27) mm, 4-4.5 times longer than wide, glabrous to puberulent.

Seeds

unwinged;

seed coats smooth.

seed coat cells with surfaces pustulate or smooth.

2n

= 16.

Delphinium trolliifolium

Delphinium carolinianum

Phenology Flowering spring.
Habitat Oak woods, coastal chaparral, wet woodlands
Elevation 30-1100 m [100-3600 ft]
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CO; FL; GA; IA; IL; KS; KY; LA; MN; MO; MS; NC; ND; NE; OK; SC; SD; TN; TX; WI; MB; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Delphinium trolliifolium occurs in the northern Coast Range of California, the Columbia River Valley to just east of Mt. Hood, and the Willamette Valley of Oregon upstream to Lane County. California plants differ somewhat from Oregon plants in pubescence patterns and habitat preferences. Further study may show that two entities are involved here.

Hybrids between Delphinium trolliifolium and D. decorum, D. menziesii subsp. pallidum (D. ×pavonaceum Ewan, Peacock larkspur), D. nudicaule, D. nuttallianum, and D. nuttallii are known. Delphinium trolliifolium is likely to be confused only with D. bakeri. Refer to discussion under that species for differences.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves absent at anthesis, cauline leaves divided into many narrow (less than 2 mm wide) segments, blade not distinctly 3-parted; distalmost petiole less than 5 mm; sepals usually blue or purple (rarely white).
subsp. carolinianum
1. Basal leaves usually present at anthesis, and/or cauline leaf lobes usually wider than 2 mm, blade often distinctly 3-parted or more; distalmost petiole more than 5 mm; sepals blue or white.
→ 2
2. Leaf blade distinctly 3-parted with few additional divisions; sepals blue to white; roots usually ± vertical, often without major branches.
subsp. vimineum
2. Leaf blade with 3–5 or more major divisions, each further divided into segments; sepals white to very pale blue; roots ±horizontal with several major branches.
→ 3
3. Stems usually less than 45 cm; in thin soils over limestone in clearings of deciduous woods; leaf blade with 3 major divisions; e of Mississippi River.
subsp. calciphilum
3. Stems usually more than 45 cm; in deeper soils in grasslands; leaf blade with 5 or more major divisions; w of Mississippi River.
subsp. virescens
Source FNA vol. 3. Treatment author: Michael J. Warnock. FNA vol. 3. Treatment author: Michael J. Warnock.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Virescens
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nudicaule, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Subordinate taxa
D. carolinianum subsp. calciphilum, D. carolinianum subsp. carolinianum, D. carolinianum subsp. vimineum, D. carolinianum subsp. virescens
Name authority A. Gray: Proc. Amer. Acad. Arts 8: 375. (1872) Walter: Fl. Carol., 155. (1788)
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