Delphinium nuttallianum
Delphinium gypsophilum
dwarf, meadow, Nuttall's larkspur, or Sonne's larkspur, slim, thin-petal larkspur, two-lobe larkspur, upland larkspur
gypsum-loving larkspur, Pinoche Creek larkspur
unbranched, 10-40(-70) cm;
base reddish, pubescence variable.
(30-)60-100(-150) cm;
base usually reddish, glabrous, glaucous.
blade round, 1-6 × 2-12 cm, nearly glabrous; ultimate lobes 5-21, 5 or more extending more than 3/5 distance to petiole, width 1-7(-14) mm (basal), 0.5-6 mm (cauline), widest at middle or in proximal 1/2.
blade round to pentagonal, 1.5-6 × 2-12 cm, nearly glabrous; ultimate lobes 3-12, width 3-24 mm (basal), 1-8 mm (cauline).
4-18(-48)-flowered, at least 2 times as long as wide;
pedicel 0.8-6 cm, pubescence variable;
bracteoles 3-8(-18) mm from flowers, green to blue, linear, 3-7 mm, pubescence variable.
15-30(-64)-flowered, cylindric;
pedicel spreading, (0.5-)1.5-3.5 cm, glabrous;
bracteoles 2-6 mm from flowers, green, linear, 2-5 mm, glabrous.
sepals usually bluish purple, rarely white to pink, puberulent, lateral sepals reflexed or spreading, 8-21 × 3-10 mm, spurs decurved to straight, ascending 20-60° above horizontal, 8-23 mm;
lower petal blades elevated, exposing stamens, blue to purple, except sometimes in white-flowered plants, 4-11 mm, clefts 2-5 mm;
hairs mostly on inner lobes below junction of blade and claw, white, rarely yellow.
sepals rarely reflexed, white to pink, nearly glabrous, lateral sepals spreading, 7-19 × 3-10 mm, spurs straight to upcurved, ascending 30-45° above horizontal, 7-15 mm;
lower petal blades elevated, exposing stamens, 3-8 mm, clefts 1-4 mm;
hairs centered near base of cleft, ± evenly distributed, white.
7-22 mm, 3.5-5 times longer than wide, glabrous to puberulent.
9-18 mm, 2.5-3.2 times longer than wide, puberulent.
winged or not;
seed coat cell surfaces smooth or roughened, blunt hairs absent.
seed coat cells brick-shaped, cell margins undulate, surfaces roughened.
= 16.
Delphinium nuttallianum
Delphinium gypsophilum
Delphinium nuttallianum represents an extremely difficult complex, with many variations in a number of morphologic traits. The complex has been and continues to be a major source of confusion for identification of Delphinium in North America. Type specimens of D. nuttallianum represent plants growing under dry conditions in open areas. These are typically found at 1200-2000 m in sage scrub or lower montane forest. Delphinium nuttallianum may be confused with D. andersonii, D. antoninum, D. depauperatum, D. gracilentum, and two subspecies of D. patens (subsp. patens and subsp. montanum). Features that may be used to separate D. nuttallianum from the first four, are enumerated under the respective species discussions. From D. patens subsp. patens, D. nuttallianum may be distinguished by its narrower leaf lobes, larger fruits, and more compact inflorescence. The frequent presence of glandular hairs in the inflorescence of D. patens subsp. montanum, contrasted with their absence in D. nuttallianum, will separate these taxa. Dwarfed plants of D. polycladon may be confused with D. nuttallianum. The latter, however may be distinguished by its ringed seeds, and it does not have prominent buds or sigmoid pedicel.
Hybrids have been seen between Delphinium nuttallianum and D. andersonii, D. depauperatum (D. ×burkei Greene), D. distichum (D. ×diversicolor Rydberg), D. nudicaule, and D. polycladon.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies 2 (2 in the flora).
Delphinium gypsophilum is sometimes confused with D. hesperium subsp. pallescens, D. recurvatum, and the white-flowered phases of D. hansenii subsp. kernense. The echinate seeds and long-haired petioles of D. hansenii immediately distinguish it from D. gypsophilum, which has neither.
Delphinium gypsophilum is related, and similar in many respects, to D. recurvatum. The two may be distinguished morphologically by their sepals. Delphinium recurvatum has reflexed, blue sepals; those of D. gypsophilum are spreading and white, although they may change to light blue when dry. Plants of D. recurvatum normally are less than 60 cm; those of D. gypsophilum are usually more than 60 cm. Ecologically, D. recurvatum occupies level ground among shrubs, typically in alkaline valley bottoms; D. gypsophilum is found on well-drained hillsides among grasses and in chaparral and oak woodland.
From Delphinium hesperium subsp. pallescens, specimens of D. gypsophilum may be separated by their much more finely dissected leaves, with less surface area, stem base usually reddish, stems frequently glaucous proximally, undulate margins of seed coat cells, and absence of striations in stem base of dried specimens. In contrast, D. hesperium subsp. pallescens has leaves less dissected, with greater surface area, stem base rarely reddish, stem not glaucous proximally, seed coat cells with straight margins, and striations present on the proximal stem of dried specimens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Lower petals 5–8 mm; lateral sepals 10 mm or more; pedicel usually more than 1 cm apart. | subsp. gypsophilum |
1. Lower petals 3–5 mm; lateral sepals 10 mm or less; pedicel usually less than 1 cm apart. | subsp. parviflorum |
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