Delphinium nuttallianum |
Delphinium cardinale |
|
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dwarf, meadow, Nuttall's larkspur, or Sonne's larkspur, slim, thin-petal larkspur, two-lobe larkspur, upland larkspur |
cardinal larkspur, cardinal or scarlet larkspur, scarlet larkspur |
|
Stems | unbranched, 10-40(-70) cm; base reddish, pubescence variable. |
(33-)50-150(-280) cm; base reddish, ± puberulent. |
Leaves | blade round, 1-6 × 2-12 cm, nearly glabrous; ultimate lobes 5-21, 5 or more extending more than 3/5 distance to petiole, width 1-7(-14) mm (basal), 0.5-6 mm (cauline), widest at middle or in proximal 1/2. |
blade round to reniform, 3-7 × 5-10 cm, nearly glabrous; ultimate lobes 0-27, width 5-40 mm (basal), 0.5-6 mm (cauline). |
Inflorescences | 4-18(-48)-flowered, at least 2 times as long as wide; pedicel 0.8-6 cm, pubescence variable; bracteoles 3-8(-18) mm from flowers, green to blue, linear, 3-7 mm, pubescence variable. |
10-40(-80)-flowered, open, narrowly pyramidal; pedicel spreading, (1-)2-5 cm, ± puberulent; bracteoles (2-)7-15(-25) mm from flowers, green, linear, 3-7 mm, glabrous to puberulent. |
Flowers | sepals usually bluish purple, rarely white to pink, puberulent, lateral sepals reflexed or spreading, 8-21 × 3-10 mm, spurs decurved to straight, ascending 20-60° above horizontal, 8-23 mm; lower petal blades elevated, exposing stamens, blue to purple, except sometimes in white-flowered plants, 4-11 mm, clefts 2-5 mm; hairs mostly on inner lobes below junction of blade and claw, white, rarely yellow. |
sepals red, glabrous, lateral sepals forward pointing, 11-15 × 5-8 mm, spurs straight, stout, slightly ascending, 15-24 mm; lower petal blades nearly coplanar with claw, exposing stamens, 2-5 mm, clefts 0.5-1.5 mm; hairs centered at base of cleft, short, sparse, yellow. |
Fruits | 7-22 mm, 3.5-5 times longer than wide, glabrous to puberulent. |
erect, 9-18 mm, 2.5-4 times longer than wide, glabrous. |
Seeds | winged or not; seed coat cell surfaces smooth or roughened, blunt hairs absent. |
unwinged; seed coat cells with margins undulate, surfaces roughened. |
2n | = 16. |
= 16. |
Delphinium nuttallianum |
Delphinium cardinale |
|
Phenology | Flowering spring (-early summer). | Flowering spring–early summer. |
Habitat | Open coniferous woods, grassy sage scrub, meadow edges and well drained streamsides (generally not in very wet sites) | Slopes (often unstable) in chaparral |
Elevation | 300-3500 m (1000-11500 ft) | 50-1500 m (200-4900 ft) |
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
|
CA; Mexico (Baja California, Baja California Sur)
|
Discussion | Delphinium nuttallianum represents an extremely difficult complex, with many variations in a number of morphologic traits. The complex has been and continues to be a major source of confusion for identification of Delphinium in North America. Type specimens of D. nuttallianum represent plants growing under dry conditions in open areas. These are typically found at 1200-2000 m in sage scrub or lower montane forest. Delphinium nuttallianum may be confused with D. andersonii, D. antoninum, D. depauperatum, D. gracilentum, and two subspecies of D. patens (subsp. patens and subsp. montanum). Features that may be used to separate D. nuttallianum from the first four, are enumerated under the respective species discussions. From D. patens subsp. patens, D. nuttallianum may be distinguished by its narrower leaf lobes, larger fruits, and more compact inflorescence. The frequent presence of glandular hairs in the inflorescence of D. patens subsp. montanum, contrasted with their absence in D. nuttallianum, will separate these taxa. Dwarfed plants of D. polycladon may be confused with D. nuttallianum. The latter, however may be distinguished by its ringed seeds, and it does not have prominent buds or sigmoid pedicel. Hybrids have been seen between Delphinium nuttallianum and D. andersonii, D. depauperatum (D. ×burkei Greene), D. distichum (D. ×diversicolor Rydberg), D. nudicaule, and D. polycladon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hybrids between Delphinium cardinale and D. parryi have been named D. ×inflexum Davidson. Because of horticultural interest in red-flowered delphiniums, garden hybrids have been made with D. elatum, D. hesperium, D. hutchinsoniae, D. nudicaule, D. parishii, D. penardii, D. scopulorum, D. tatsienense Franchet, D. uliginosum, and D. zalil Aitchison & Hemsley, although D. cardinale does not grow with any of these in the wild. Plants of Delphinium cardinale are quite variable in size, leaf distribution, and pubescence, resulting in considerable differences between, and sometimes within, populations. No patterns could be seen, however, to justify recognition of separate taxa within D. cardinale. Populations farther south (in Baja California, Mexico) may represent a distinct entity; they require further study. The only possible confusion between Delphinium cardinale (seeds not ringed, fruits erect, grows in relatively dry sites) and another taxon might occur with Delphinium nudicaule (seeds ringed, fruits spreading, grows in moist habitats). The two are separated geographically and phenologically (although D. cardinale may begin flowering in southern California before D. nudicaule has finished in northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 3. | FNA vol. 3. |
Parent taxa | Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Grumosa | Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Wislizenana |
Sibling taxa | ||
Synonyms | D. pauciflorum, D. nuttallianum var. fulvum, D. nuttallianum var. levicaule, D. sonnei | |
Name authority | Pritzel: in Walpers, Repert. Bot. Syst. 1: 744. (1842) | Hooker: Bot. Mag., plate 4887. (1855) |
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