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Delphinium nudicaule

canyon delphinium, canyon larkspur, orange larkspur, red larkspur, red or orange larkspur, scarlet larkspur

delphinium, larkspur

Habit Herbs, perennial, from fasciculate roots or rhizomes.
Stems

(15-)20-50(-125) cm;

base reddish, glabrous.

Leaves

blade round to pentagonal, 2-6 × 3-10 cm; ultimate lobes 3-12, width 5-40 mm (basal), 2-20 mm (cauline).

blade deeply palmately divided, round to pentagonal or reniform, margins entire or lobes apically crenate or lacerate, lobes of basal blades wider and fewer than those of cauline blades.

Inflorescences

5-20(-69)-flowered;

pedicel (1.5-)2-6(-8) cm, glabrous to glandular-pubescent;

bracteoles 14-20(-30) mm from flowers, green to red, linear, 2-4(-9) mm, glabrous to puberulent.

terminal, 2-100(-more)-flowered racemes (occasionally branched, thus technically panicles), 5-40 cm or more;

bracts subtending inflorescence branches;

pedicels present or absent;

bracteoles (on pedicels) subopposite-subalternate, not forming involucre.

Flowers

sepals scarlet to reddish orange, rarely dull yellow, glabrous, lateral sepals forward-pointing to form pseudotube, (6-)8-13(-16) × 3-6 mm, spurs straight, slightly ascending, (12-)18-27(-34) mm;

lower petal blades elevated, exposing stamens, 2-3 mm, clefts 0.5-1 mm;

hairs sparse, evenly dispersed, yellow.

bisexual, bilaterally symmetric;

sepals not persistent in fruit, 5;

upper sepal 1, spurred, 8-24 mm;

lateral sepals 2, ± ovate to elliptic, 8-18 mm;

lower sepals 2, similar to lateral sepals;

upper petals 2, spurred, enclosed in upper sepal, nectary inside tip of spur;

lower petals 2, plane, ± ovate, ± 2-lobed, clawed, 2-12 mm, nectary absent;

stamens 25-40;

filaments with base expanded;

staminodes absent between stamens and pistils;

pistils 3(-5), simple;

ovules 8-20 per pistil;

style present.

Fruits

13-26 mm, 3.5-4.5 times longer than wide, glabrous.

follicles, aggregate, sessile, ± curved-cylindric, sides prominently veined or not;

beak terminal, straight, 2-4 mm.

Seeds

unwinged or sometimes slightly wing-margined;

seed coat cells with surfaces smooth.

dark brown to black (often appearing white because of air in seed coat cells), rectangular to pyramidal, often ± rough surfaced.

x

= 8.

2n

= 16.

Delphinium nudicaule

Delphinium

Phenology Flowering late winter–early summer.
Habitat Moist talus, cliff faces
Elevation 0-2600 m (0-8500 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from USDA
n temperate and arctic subtropical and; in Eastern Hemisphere; tropical mountains (s of equator in Africa)
[BONAP county map]
Discussion

Delphinium nudicaule hybridizes with most other taxa of Delphinium that it encounters. Apparent hybrids involving D. nudicaule, and seen by the author (either afield or as specimens), include D. andersonii, D. antoninum, D. decorum, D. luteum, D. nuttallianum, D. patens, and D. trolliifolium. In addition, garden-grown plants have been hybridized with D. cardinale, D. elatum, D. menziesii, D. parishii, D. penardii, D. tatsienense Franchet, D. triste Fischer ex de Candolle, and D. uliginosum; D. nudicaule does not naturally occur with these species. Delphinium nudicaule is one of the earliest larkspurs to flower in any given locality. Douglas's type collection of D. nudicaule represents plants (synonyms D. sarcophyllum Hooker & Arnott and D. peltatum Hooker, an invalid name) grown under very moist conditions, probably quite near the ocean. The type specimen of D. armeniacum A. Heller represents plants grown under unusually dry conditions.

The Mendocino Indians consider Delphinium nudicaule a narcotic (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 300 (61 in the flora).

Three Eurasian species of Delphinium–D. elatum Linnaeus, D. grandiflorum Linnaeus, and D. tatsienense Franchet–have been commonly cultivated in North America. Of the nonnative taxa, only D. elatum is sporadically naturalized, as far as is known. Isolating mechanisms in Delphinium appear to be primarily ecological, geographic, and/or temporal. Where these distinctions are disrupted, introgression often exists. Hybridization occurs regularly between certain taxa, particularly in areas of disturbance (e.g., roadcuts, drainage ditches, clearcuts). The more common and easily recognized hybrids are included in the key.

Many names have been misapplied in Delphinium. The few misapplied names mentioned in discussions below refer to relatively widespread problems.

Unless otherwise noted, the key and descriptions refer to fresh material. Some features may be significantly altered by pressing; they can, however, usually be determined with a certain amount of effort and experience.

In the descriptions, "base of cleft" refers to the point where the cleft or sinus reaches most deeply into the petal blade.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lower petal blades less than 1/5 length of lateral sepals; sepals never red or yellow.
Sect. Elatopsis
1. Lower petal blades more than 1/5 length of lateral sepals; sepals blue, purple, white, red, or yellow.
Sect. Diedropetala
Source FNA vol. 3. FNA vol. 3. Author: Michael J. Warnock.
Parent taxa Ranunculaceae > Delphinium > sect. Diedropetala > subsect. Bicoloria Ranunculaceae
Sibling taxa
D. alabamicum, D. alpestre, D. andersonii, D. andesicola, D. antoninum, D. bakeri, D. barbeyi, D. basalticum, D. bicolor, D. brachycentrum, D. californicum, D. cardinale, D. carolinianum, D. decorum, D. depauperatum, D. distichum, D. elatum, D. exaltatum, D. geraniifolium, D. geyeri, D. glareosum, D. glaucescens, D. glaucum, D. gracilentum, D. gypsophilum, D. hansenii, D. hesperium, D. hutchinsoniae, D. inopinum, D. lineapetalum, D. luteum, D. madrense, D. menziesii, D. multiplex, D. newtonianum, D. novomexicanum, D. nuttallianum, D. nuttallii, D. parishii, D. parryi, D. patens, D. polycladon, D. purpusii, D. ramosum, D. recurvatum, D. robustum, D. sapellonis, D. scaposum, D. scopulorum, D. stachydeum, D. sutherlandii, D. treleasei, D. tricorne, D. trolliifolium, D. uliginosum, D. umbraculorum, D. variegatum, D. viridescens, D. wootonii, D. xantholeucum
Subordinate taxa
Sect. Diedropetala, Sect. Elatopsis
Synonyms D. armeniacum
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 33. (1838) Linnaeus: Sp. Pl. 1: 530. 175: Gen. Pl. ed 5, 236. (1754)
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